(2005).
Center for Invasive Plant Management. (2008).
Davis, Mark, et al. “Fluctuating Resources in Plant Communities: A General Theory of Invasibility.” Journal of Ecology 88 (2000): 528-534.
Dillenburg, Lucia, et al. “Effects of Vine Competition on Availability of Light, Water, and Nitrogen to a Tree Host (Liquidambar Styraciflua).” American Journal of Botany 80 (1993): 244-252.
Fox, Alison M. and Doria R. Gordon. “Criteria for Listing Invasive Plants.” Weed Technology 18 (2004): 1309-1313.
Jefferson, Lara, et al. “Implementing Invasive Screening Procedures: The Chicago Botanical Model.” Weed Technology 18 (2004): 1434-1440.
MacDonald, Greg, et al. “Japanese Honeysuckle.” Center for Aquatic and Invasive Plants (2008).
National Invasive Species Council. (2008).
Nyboer, Randy. “Vegetation Management Guideline: Japanese Honeysuckle.” (1988).
Oklahoma Invasive Plant Council. (2008).
Powell, Mark R. “Risk Assessment for Invasive Plant Species.” Weed Technology 18 (2004): 1305-1308.
Regehr, David L. and David R. Frey. “Selective Control of Japanese Honeysuckle (Lonicera japonica).” Weed Technology 2 (1988): 139-143.
Richardson, David M., et al. “Naturalization and Invasion of Alien Plants: Concepts and Definitions.” Diversity and Distributions 6 (2000): 93-107.
Shannon, Kim and Caryn Vaughn. “Oklahoma Natural Areas Registry: Oliver’s Woods.” Biosurvey News Fall (2003).
USDA Plants Database. “Lonicera japonica.”
Appendix
Figure 1. An aerial view of the area of interest, Oliver’s Woods, Cleveland County, Norman, OK.
List of Floral Keys Used in This Study:
-
Hunter, Carl G. Trees, shrubs and vines of Arkansas. (1989). Ozark Society Foundation
-
Little, Jr. Elbert L. Forest Trees of Oklahoma. (2002). State Dept. of Agriculture
-
Smith, Edwin B. Keys to the Flora of Arkansas. (1994). The University of Arkansas Press
Daubenmeyer Scale
|
Value
|
% Coverage
|
0
|
0
|
1
|
0.1-1.0
|
2
|
1.0-5.0
|
3
|
5.0-25.0
|
4
|
25.0-50.0
|
5
|
50.0-75.0
|
6
|
75.0-90.0
|
7
|
90.0-100.0
|
Figure 2. The percentage of canopy openness for plots.
Figure 3. Soil pH for plots.
Figure 4. Soil Nitrogen for plots.
Figure 5. Soil Phosphorous for plots.
Soil Moisture (g)
|
H1
|
0.023
|
R1
|
0.114
|
H2
|
0.059
|
R2
|
0.113
|
H3
|
0.039
|
R3
|
0.234
|
H4
|
0.000
|
R4
|
0.422
|
H5
|
0.009
|
R5
|
0.129
|
H6
|
0.012
|
R6
|
0.143
|
H7
|
0.012
|
R7
|
0.137
|
H8
|
0.000
|
R8
|
0.168
|
H9
|
0.063
|
R9
|
0.130
|
H10
|
0.083
|
R10
|
0.225
|
H11
|
0.019
|
R11
|
0.113
|
H12
|
0.089
|
R12
|
0.164
|
H13
|
0.102
|
R13
|
0.153
|
H14
|
0.104
|
R14
|
0.152
|
H15
|
0.146
|
R15
|
0.281
|
H16
|
0.107
|
R16
|
0.145
|
H17
|
0.024
|
R17
|
0.278
|
H18
|
0.018
|
R18
|
0.116
|
H19
|
0.088
|
R19
|
0.147
|
H20
|
0.084
|
R20
|
0.202
|
Table 1. Soil moisture content for plots.
Plot
|
Nearest Tree
|
Distance to Tree (m)
|
Tree Diameter (m)
|
Clump Circumference (m)
|
H1
|
Acer negundo
|
0.40
|
1.00
|
4.09
|
H2
|
Juglans nigra
|
0.35
|
0.73
|
3.27
|
H3
|
Juglans nigra
|
0.85
|
2.34
|
8.58
|
H4
|
Celtis occidentalis
|
0.94
|
2.24
|
6.97
|
H5
|
Juglans nigra
|
0.59
|
1.25
|
8.19
|
H6
|
Ligustrum vulgare
|
0.27
|
0.99
|
9.07
|
H7
|
Cercis canadensis
|
0.40
|
0.39
|
7.08
|
H8
|
Tilia americana
|
0.11
|
1.40
|
6.11
|
H9
|
Carya illinoensis
|
0.63
|
1.01
|
6.01
|
H10
|
Carya illinoensis
|
0.41
|
0.79
|
7.32
|
H11
|
Carya illinoensis
|
0.63
|
0.91
|
1.50
|
H12
|
Celtis occidentalis
|
0.77
|
2.40
|
7.39
|
H13
|
Cercis canadensis
|
0.53
|
0.92
|
5.97
|
H14
|
Juglans nigra
|
0.84
|
1.06
|
3.16
|
H15
|
Juniperus virginiana
|
0.37
|
1.52
|
2.61
|
H16
|
Celtis laevigata
|
0.34
|
1.48
|
11.80
|
H17
|
Quercus macrocarpa
|
0.12
|
0.26
|
7.95
|
H18
|
Ulmus sp.
|
0.21
|
0.35
|
10.45
|
H19
|
Celtis occidentalis
|
0.20
|
1.40
|
9.63
|
H20
|
Quercus macrocarpa
|
0.11
|
1.92
|
20.00
|
Table 2. Tree nearest to clump of honeysuckle id, distance from clump, and diameter. Also honeysuckle clump circumference.
Plot
|
Nearest Tree
|
Distance to Tree (m)
|
Tree Diameter (m)
|
R1
|
Juglans nigra
|
0.40
|
1.20
|
R2
|
Juglans nigra
|
0.10
|
1.28
|
R3
|
Cercis canadensis
|
0.71
|
2.00
|
R4
|
Cercis canadensis
|
0.55
|
2.93
|
R5
|
Cercis canadensis
|
1.13
|
2.52
|
R6
|
Juglans nigra
|
0.62
|
2.07
|
R7
|
Fraxinus pennsylvanica
|
0.90
|
1.01
|
R8
|
Fraxinus pennsylvanica
|
0.44
|
0.38
|
R9
|
Carya illinoinensis
|
0.92
|
1.33
|
R10
|
Juniperus virginiana
|
0.59
|
2.09
|
R11
|
Carya illinoensis
|
0.22
|
2.19
|
R12
|
Cercis canadensis
|
0.74
|
1.51
|
R13
|
Celtis occidentalis
|
0.36
|
2.56
|
R14
|
Juglans nigra
|
0.23
|
0.80
|
R15
|
Juniperus virginiana
|
0.28
|
1.53
|
R16
|
Juglans nigra
|
1.67
|
2.23
|
R17
|
Quercus macrocarpa
|
2.40
|
0.46
|
R18
|
Juglans nigra
|
0.32
|
3.45
|
R19
|
Celtis occidentalis
|
0.88
|
2.71
|
R20
|
Morus alba
|
0.46
|
2.53
|
Table 3. Tree nearest to clump of honeysuckle id, distance from clump, and diameter.
Coverage of Associated Species – Honeysuckle Plots
|
|
H-1
|
H-2
|
H-3
|
H-4
|
H-5
|
H-6
|
H-7
|
H-8
|
H-9
|
H-10
|
H-11
|
H-12
|
H-13
|
H-14
|
H-15
|
H-16
|
H-17
|
H-18
|
H-19
|
H-20
|
Apiaceae sp.
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
2
|
0
|
1
|
3
|
0
|
1
|
3
|
1
|
0
|
1
|
2
|
1
|
Aster sp.
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Bidens bipinnata
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Brassicaceae sp.
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
3
|
0
|
3
|
2
|
0
|
1
|
1
|
1
|
Bromus sp.
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
4
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Carex sp.
|
3
|
4
|
0
|
0
|
1
|
0
|
1
|
0
|
2
|
3
|
3
|
0
|
3
|
3
|
3
|
2
|
3
|
3
|
2
|
1
|
Carya illinoensis
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Carya illinoensis seedling
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
Celtis occidentalis
|
0
|
0
|
0
|
0
|
0
|
0
|
4
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Celtis occidentalis seedling
|
0
|
0
|
4
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
4
|
3
|
0
|
0
|
2
|
0
|
2
|
Cercis canadensis seedling
|
2
|
0
|
0
|
3
|
2
|
3
|
0
|
0
|
0
|
0
|
4
|
3
|
0
|
0
|
1
|
0
|
0
|
1
|
0
|
0
|
Elephantopus carolinianus
|
0
|
3
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Eupatorium
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
0
|
0
|
Geum canadense
|
1
|
0
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
3
|
0
|
0
|
1
|
1
|
2
|
1
|
1
|
0
|
2
|
Gallium sp.
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
1
|
0
|
0
|
0
|
Galium triflorium
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Ivy cultivar-periwinkle
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
4
|
Ligustrum vulgare
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Lonicera japonica
|
4
|
5
|
3
|
6
|
6
|
3
|
4
|
6
|
6
|
2
|
3
|
3
|
3
|
5
|
3
|
6
|
7
|
3
|
6
|
5
|
Morus rubra sapling
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Opuntia humifusa
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Panicum sp.
|
0
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
3
|
0
|
1
|
0
|
0
|
3
|
0
|
0
|
Parthenocissus quinquefolia
|
0
|
0
|
0
|
0
|
1
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
0
|
0
|
1
|
0
|
Poaceae sp.
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Polygonum sp.
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
Rhus copallina
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Rosa multiflora
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Smilax bona-nox
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
3
|
0
|
0
|
0
|
0
|
Smilax tamnoides var. hispida
|
0
|
0
|
2
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
0
|
2
|
0
|
0
|
Symphoricarpos orbiculatus
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
0
|
3
|
3
|
6
|
2
|
0
|
0
|
5
|
0
|
3
|
3
|
3
|
Tilia americana
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Toxicodendron radicans
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
Ulmus sp.
|
0
|
0
|
2
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
Vitis sp.
|
0
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
1
|
1
|
0
|
0
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
|