Ecological Dynamics of the Site
This ecological site occurs on weathered ash soils in warm, very wet regions of North Kohala district of the Island of Hawai`i. Plant communities evolved without the presence of large mammals or the regular occurrence of fires. Much of the original plant community is still relatively intact, but has been impacted by weed invasion and damage from feral animals.
State and transition diagram
State 1 – Native Dwarf Forest
Plant Community 1
This state represents the Historic Climax Plant Community. The general aspect is a thicket of low stature trees and shrubs with native grasses in openings; the soil surface is covered by a deep layer of sphagnum moss. The tallest plants, primarily ohia and olapa, typically range from 4 to 15 feet tall. In some areas, the overstory may be as short as 3 feet or as tall as 30 feet. Where the tree canopy is tallest, the plant community has a more open understory, grading into the appearance of the plant community found in Ecological Site F164XY500HI-Mid-Stature Ohia-Olapa Forest. Tree and shrub trunks and branches are covered by a heavy growth of mosses, liverworts, and small ferns.
Pathways from this state/plant community
To State 2, Weedy Dwarf Forest, via “A&D”:
A = weed invasion; D = ungulate disturbance.
Native Dwarf Forest degrades to Weedy Dwarf Forest by gradual replacement of the understory by alien herbs and small trees that outcompete native understory species. This process is accelerated by ungulate foraging that disturbs the soil surface and directly destroys native plants and prevents their regeneration.
Plant species listed in the following tables have been observed in the course of field work or are derived from reliable records.
Abbreviations:
Origin: n = native (endemic or indigenous); a = alien (introduced by humans).
Type: t = tree; tf = tree fern; s = shrub; h = herb (forb); v = vine; f = fern; g = grasslike (grasses, sedges, rushes).
Composite representation of State 1, Plant Community 1, Native Dwarf Forest.
Scientific name
|
%Canopy cover by height class (ft)
|
Total Cover
|
Local
common name
|
NRCS
common name
|
Origin
|
Type
|
NRCS
Code
|
0.1 -
2
|
2.1 -
4.5
|
4.6 -
13
|
13.1 -
40
|
40.1 -
80
|
80.1 -
120
|
Metrosideros polymorpha
|
1
|
50
|
20
|
|
|
|
70
|
'ohi'a lehua
|
'ohi'a lehua
|
n
|
t
|
MEPO5
|
Cheirodendron trigynum
|
1
|
5
|
5
|
|
|
|
10
|
olapa
|
olapalapa
|
n
|
t
|
CHTR2
|
Myrsine lessertiana
|
1
|
1
|
|
|
|
|
1
|
kolea lau nui
|
kolea lau nui
|
n
|
t
|
MYLE2
|
Myrsine sandwicensis
|
1
|
5
|
1
|
|
|
|
5
|
kolea lau li`i
|
kolea lau li`i
|
n
|
t
|
MYSA2
|
Melicope sp.
|
tr
|
1
|
1
|
|
|
|
1
|
alani
|
melicope
|
n
|
t
|
MELIC3
|
Ilex anomala
|
tr
|
1
|
|
|
|
|
1
|
kawa`u
|
Hawai`i holly
|
n
|
t
|
ILAN
|
Hedyotis terminalis
|
tr
|
1
|
|
|
|
|
1
|
manono
|
variable starviolet
|
n
|
t
|
HETE21
|
Clermontia drepanomorpha
|
tr
|
1
|
1
|
|
|
|
1
|
`oha wai
|
Kohala mountain clermontia
|
n
|
t
|
CLDR2
|
Cyanea tritomantha
|
tr
|
1
|
|
|
|
|
1
|
`aku
|
`aku`aku
|
n
|
t
|
CYTR6
|
Pritchardia lanigera
|
tr
|
tr
|
1
|
1
|
|
|
1
|
loulu
|
lou`ulu
|
n
|
t
|
PRLA4
|
Cibotium glaucum
|
1
|
1
|
1
|
|
|
|
1
|
hapu`u
|
hapu`u
|
n
|
tf
|
CIGL
|
Cibotium menziesii
|
1
|
1
|
tr
|
|
|
|
1
|
hapu`u `i`i
|
hapu`u li
|
n
|
tf
|
CIME8
|
Cibotium chamissoi
|
tr
|
1
|
tr
|
|
|
|
1
|
hapu`u
|
Chamisso's manfern
|
n
|
tf
|
CICH
|
Labordia hedyosmifolia
|
tr
|
1
|
|
|
|
|
1
|
kamakahala
|
bog labordia
|
n
|
s
|
LAHE2
|
Broussaisia arguta
|
tr
|
1
|
tr
|
|
|
|
1
|
kanawao
|
kanawao
|
n
|
s
|
BRAR6
|
Vaccinium calycinum
|
1
|
30
|
|
|
|
|
30
|
ohelo
|
ohelo kau la`u
|
n
|
s
|
VACA8
|
Vaccinium dentatum
|
1
|
|
|
|
|
|
1
|
ohelo
|
ohelo
|
n
|
s
|
VADE2
|
Styphelia tameiameiae
|
1
|
1
|
|
|
|
|
1
|
pukiawe
|
pukiawe
|
n
|
s
|
STTA
|
Clermontia calophylla
|
1
|
1
|
|
|
|
|
1
|
`oha wai
|
lava clermontia
|
n
|
s
|
CLCA5
|
Clermontia waimeae
|
1
|
1
|
|
|
|
|
1
|
`oha wai
|
swampforest clermontia
|
n
|
s
|
CLWA2
|
Trematolobelia grandifolia
|
1
|
1
|
tr
|
|
|
|
1
|
|
largeflower false lobelia
|
n
|
s
|
TRGR8
|
Dubautia plantaginea
|
tr
|
tr
|
tr
|
|
|
|
tr
|
na`ena`e
|
plantainleaf dubautia
|
n
|
s
|
DUPL
|
Eurya sandwicensis
|
tr
|
tr
|
|
|
|
|
tr
|
anini
|
anini
|
n
|
s
|
EUSA6
|
Coprosma ochracea
|
tr
|
1
|
|
|
|
|
1
|
pilo
|
Maui mirrorplant
|
n
|
s
|
COOC3
|
Viola maviensis
|
tr
|
|
|
|
|
|
tr
|
|
Hawai`i bog violet
|
n
|
s
|
VIMA6
|
Astelia menziesiana
|
1
|
|
|
|
|
|
1
|
kaluaha
|
pua'akuhinia
|
n
|
h
|
ASME4
|
Peperomia sp.
|
1
|
|
|
|
|
|
1
|
`ala`ala wai nui
|
peperomia
|
n
|
h
|
PEPER
|
Phyllostegia floribunda
|
tr
|
|
|
|
|
|
tr
|
|
Hawai`i phyllostegia
|
n
|
h
|
PHFL6
|
Phyllostegia vestita
|
tr
|
|
|
|
|
|
tr
|
|
streambed phyllostegia
|
n
|
h
|
PHVE4
|
Dubautia scabra
|
tr
|
|
|
|
|
|
tr
|
na`ena`e
|
rough dubautia
|
n
|
h
|
DUSC
|
Coprosma granadensis
|
tr
|
|
|
|
|
|
tr
|
makole
|
makole
|
n
|
h
|
COGR23
|
Alyxia oliviformis
|
1
|
|
|
|
|
|
1
|
maile
|
maile
|
n
|
v
|
ALOL2
|
Smilax melastomifolia
|
1
|
|
|
|
|
|
1
|
hoi kuahiwi
|
Hawai`i greenbrier
|
n
|
v
|
SMME
|
Freycinetia arborea
|
1
|
|
|
|
|
|
1
|
`ie`ie
|
`ie`ie
|
n
|
v
|
FRAR
|
Stenogyne cranwelliae
|
1
|
|
|
|
|
|
1
|
|
bog stenogyne
|
n
|
v
|
STCA9
|
Stenogyne calaminthoides
|
1
|
|
|
|
|
|
1
|
|
|
|
|
|
Phyllostegia warshaueri
|
tr
|
|
|
|
|
|
tr
|
|
Laupahoehoe phyllostegia
|
n
|
v
|
PHWA3
|
Sadleria sp.
|
1
|
1
|
|
|
|
|
1
|
`ama`u
|
sadleria
|
n
|
f
|
SADLE
|
Elaphoglossum crassifolium
|
1
|
|
|
|
|
|
1
|
stag's tongue, `ekaha
|
royal tonguefern
|
n
|
f
|
ELCR2
|
Elaphoglossum paleaceum
|
1
|
|
|
|
|
|
1
|
maku`e
|
ekaha
|
n
|
f
|
ELHI3
|
Athyrium microphyllum
|
1
|
|
|
|
|
|
1
|
`akolea
|
akolea
|
n
|
f
|
ATMI
|
Asplenium sp.
|
1
|
|
|
|
|
|
1
|
|
spleenwort
|
n
|
f
|
ASPLE
|
Lycopodiella cernua
|
1
|
|
|
|
|
|
1
|
pakahakaha
|
weeping fern
|
n
|
f
|
LETH6
|
Adenophorus tamariscinus
|
1
|
|
|
|
|
|
1
|
wahini noho mauna
|
wahini noho mauna
|
n
|
f
|
ADTA
|
Adenophorus pinnatifidus
|
1
|
|
|
|
|
|
1
|
|
graceful kihifern
|
n
|
f
|
ADPI
|
Dryopteris wallichiana
|
1
|
|
|
|
|
|
1
|
`i`o nui
|
alpine woodfern
|
n
|
f
|
DRWA
|
Sticherus owhyhensis
|
1
|
|
|
|
|
|
1
|
uluhe
|
Hawai`i umbrella fern
|
n
|
f
|
STOW
|
|