Appendix 1. Species included in the molecular analysis with voucher information and GenBank accession numbers (ITS1;
ITS2; rpS4-trnT-trnL). For all species of Cousinia the subgeneric classification is given. An asterisk (*) indicates sequences previously published.
Hypacanthium echinopifolium (Bornm.) Juz., Kyrgyzstan: Sine loc., Iljin s.n. (LE), AY373746*; AY373713*; EU661125. Jurinea
albicaulis Bunge, Greece: Makedonia, Halkidiki, between Arnea and Paleokastro, Susanna 1957 & al. (BC), AY373747*; AY373714*;
EU661092. Jurinea humilis DC., Spain: Huesca, Plan, La Barana, P. Montserrat s.n. (JACA), L35868*; EU661093. Saussurea
elegans Ledeb., Kazakhstan: Aksu-Dzabagly reservation, 1,800 m, Susanna 2179 & al. (BC), AY373750*; AY373717*; EU661090.
Saussurea maximowiczii Herder, Japan: Ofuna Botanical Garden, AY373751*; AY373718*; EU661091. Schmalhausenia nidulans
(Regel) Petrak, Kazakhstan: Almatinskaya oblast, Alatau mt., above Almaty, Susanna 2088 & al. (BC), AY373752*; AY373719*;
EU661124. Tiarocarpus hymenostephanus Rech. f., Afghanistan: W, Herat, Band-e Farsi, 10 SW Farsi, 3,200 m, Freitag 6841
(KAS), EU923831; EU923945; EU661139. Tiarocarpus neubaueri (Rech. f.) Rech. f., Afghanistan: C, Bamian, Band-i Amir,
Hochfächen, 3,000 m, Dieterle 896 (M), EU923775; EU923889; EU661140. Arctium lappa L., Iran: NNE, Khorassan: 25 SW
Bodjnourd, 1,500 m, Mehregan 140 (MJG), EU923773; EU923887; EU661123. Arctium leiospermum Juz. & C. Serg., Kazakhstan:
Zambylskaya oblast, Kurdai pass, 900 m, Susanna 2154 & al. (BC), AY373720*; AY373687*; EU661121. Arctium minus (Hill.)
Bernh., Belgium: Lovaina Botanical Garden, AF19049*; AF19103*; EU661122. Cousinia oopoda Juz. (subg. Cousinia sect. Actinia
Bunge), Tajikistan: South Tajikistan, west mountainside, Khodzha-Kazjyan range, Koy-Pjez-Tau mt., limestone outcrops, 1,240 m,
Kinzikaeva 3585 (LE), EU923839; EU923953; EU661232. Cousinia tashkurghanica Rech. f. (subg. Cousinia sect. Actinia Bunge),
Afghanistan: N, Mazar Sharif, Tang-e Sayyad, S of Tashkurghan, 600 m, Freitag 3905 (KAS), EU923777; EU923891; EU661234.
Cousinia oligocephala Boiss. (subg. Cousinia sect. Albidae Rech. f.), Iran: Fars, Inter Shiraz and Sepidan, Shoul pass, 2,000 m,
Mehregan 218 (MJG), EU923796; EU923910; EU661208. Cousinia chrysantha Kult. (subg. Cousinia sect. Alpinae Bunge), Kazakhstan: Shimkientskaya oblast, Aksu Dzabagly reservation, Darbassa canyon, 1,840 m, Susanna 2198 & al. (BC), AY373725*;
AY373692*; EU661136. Cousinia dipterocarpa Bornm. & Rech. f. (subg. Cousinia sect. Alpinae Bunge), Iran: Khorassan, 27 km
from Ghouchan to Bajgiran, Alam Ali pass, 1,820 m, Mehregan 167 (MJG), EU923862; EU923976; EU661155. Cousinia lasiolepis
Boiss. (subg. Cousinia sect. Alpinae Bunge), Iran: Yasoudj, Dena mts., Bizhan Pass, 3,300 m, Mehregan 202 (MJG), EU923863;
EU923977; EU661177. Cousinia multiloba DC. (subg. Cousinia sect. Alpinae Bunge), Iran: Khorassan, SW Bodjnourd, Salouk mt.,
2,200 m, Mehregan 189 (MJG), EU923835; EU923949; EU661202. Cousinia serawschanica C. Winkl. (subg. Cousinia sect. Alpinae Bunge), Tajikistan: Voru, 2,000–2,300 m, Susanna 2516 & al. (BC), EU923874; EU923988; EU661192. Cousinia splendida
C. Winkl. (subg. Cousinia sect. Alpinae Bunge), Tajikistan: Fan mountains, road above Iskandar-Kul, 2,300 m, Susanna 2507 & al.
(BC), EU923871; EU923985; EU661197. Cousinia badghysi Kult. (subg. Cousinia sect. Badghysia Tschern.), Turkmenistan:
Badghys, Eroylanduz lake, SE ¨sopki¨ Kazan, Kamelin 360 (LE), EU923872; EU923986; EU661166. Cousinia piptocephala Bunge
(subg. Cousinia sect. Badghysia Tschern.), Iran: Isfahan, 5km from Mourcheh Khort towards Natanz, 1,500 m, Mehregan 188 (MJG),
EU923801; EU923915; EU661204. Cousinia ariana Bornm. (subg. Cousinia sect. Carduncellus (Juz.) Rech. f.), Afghanistan: E,
Kabul, N Salang, 3,150 m, Freitag 3494 (KAS), EU923788; EU923902; EU661221. Cousinia buphthalmoides Regel (subg. Cousinia sect. Carduncellus (Juz.) Rech. f.), Afghanistan: Unai-Pass, E side, 3,050 m, Freitag 6268 (KAS), EU923789; EU923903;
EU661179. Cousinia tianschanica Kult. (subg. Cousinia sect. Carduncellus (Juz.) Rech. f.), Kazakhstan: Shimkientskaya oblast,
Aksu Dzabagly reservation, Aksu canyon, Susanna 2191 & al. (BC), AY373743*; AY373710*; EU661127. Cousinia dissecta Kar.
& Kir. (subg. Cousinia sect. Chrysoptera Tschern.), Kazakhstan: Almatinskaya oblast, Sugati mts., Kokpekti narrows, Susanna 2137
& al. (BC), AY373728*; AY373695*; EU661141. Cousinia oxiana Tschern. (subg. Cousinia sect. Chrysoptera Tschern.), Afghanistan: NW, Herat, 34 km N of Shindand, 1,320 m, Amin 5252 (KAS), EU923854; EU923968; EU661227. Cousinia turkmenorum
Bornm. (subg. Cousinia sect. Chrysoptera Tschern.), Iran: Khorassan, 20 km from Mashhad to Neishabour, 1,150 m, Mehregan 197
(MJG), EU923855; EU923969; EU661156. Cousinia greuteri Rech. f. (subg. Cousinia sect. Congesta Bunge), Afghanistan: N,
Robatak pass between Haibak and Puli-Khumri, 1,400 m, Tonchev 2200 (KAS), EU923783; EU923897; EU661237. Cousinia
coronata Franch. (subg. Cousinia sect. Coronophora (Juz.) Rech. f), Uzbekistan: road Tashkent–Samarkand, 10 km from the crossroads to Jizzak, 400 m, Susanna 2039 & al. (BC), AY373727*; AY373694*; EU661178. Cousinia radians Bunge (subg. Cousinia
sect. Coronophora (Juz.) Rech. f.), Iran: Khorassan, 66 km from Mashad to Kalat-Naderi, 1,000 m, Ghahreman & al. s.n. (TUH),
EU923791; EU923905; EU661217. Cousinia congesta Bunge (subg. Cousinia sect. Cousinia), Uzbekistan: between Samarkand
and Kitov, Takhta–Karachi pass, 1,600 m, Susanna 2059 & al. (BC), AY373726*; AY373693*; EU661131. Cousinia daralaghezica
Takht. (subg. Cousinia sect. Cousinia), Armenia: Vajots Dzor, Eghegnadzor distr., Eghegnadzor town, Tamanian s.n. (ERE),
EU923813; EU923927; EU661216. Cousinia intertexta Freyn & Sint. (subg. Cousinia sect. Cousinia), Turkey: Sivas, DivrigiArapkir, 20 km S Divrigi, 1,410 m, Nydegger 17154 (M), EU923817; EU923931; EU661235. Cousinia meghrica Takht. (subg.
Cousinia sect. Cousinia), Armenia: Sjunik, Meghri distr., Agarak village, Tamanian s.n. (ERE), EU923814; EU923928; EU661142.
Cousinia minkwitziae Bornm. (subg. Cousinia sect. Cousinia), Kazakhstan: Zambylskaya oblast, Talaski Alatau, 6 km W from Il
Tai, 1,000 m, Susanna 2183 & al. (BC), AY373735*; AY373702*; EU661135. Cousinia polycephala Rupr. (subg. Cousinia sect.
Cousinia), Kazakhstan: Zambylskaya oblast, 30 km from the Kurdai pass, Susanna 2161 & al. (BC), AY373738*; AY373705*;
EU661138. Cousinia syrdariensis Kult. (subg. Cousinia sect. Cousinia), Kazakhstan: Zambylskaya oblast, 30 km from the Kurdai
pass, Susanna 2159 & al. (BC), AY373741*; AY373708*; EU661133. Cousinia tenuifolia C.A. Mey. (subg. Cousinia sect. Cousinia),
Iran: Azerbaijan, 10 km from Sarab to Ardebil, 1,400 m, Mehregan 186 (MJG), EU923812; EU923926; EU661171. Cousinia urumiensis Bornm. (subg. Cousinia sect. Cousinia), Iran: Azerbaijan, Urmia, inter Silvana and Razhan, 1,650 m, Mehregan 201 (MJG),
EU923809; EU923923; EU661205. Cousinia araneosa DC. (subg. Cousinia sect. Cynaroideae Bunge), Iran: Hamedan, Alvand mt.,
10 km from Hamedan to Toiserkan, Ganjnameh Valley, 2,400 m, Mehregan 219 (MJG), EU923870; EU923984; EU6611678. Cousinia macrocephala C.A. Mey. (subg. Cousinia sect. Cynaroideae Bunge), Iran: Azerbaijan, Arasbaran, Molouk pass, 1,500 m,
Mehregan 212 (MJG), EU923852; EU923966; EU661169. Cousinia purpurea C.A. Mey. in DC. (subg. Cousinia sect. Cynaroideae
Bunge), Armenia: Ararat, near the village of Shahap, roadsides, Tamanian s.n. (ERE), AY373739*; AY373706*; EU661137. Cousinia decumbens Rech. f. (subg. Cousinia sect. Decumbentes Rech. f.), Iran: Semnan, Shahroud, mt. Shahvar, SW Slopes, 3,600 m Mehregan 110 (MJG), EU923821; EU923935; EU661157. Cousinia bungeana Regel & Schmalh. (subg. Cousinia sect. Dichotomae
Bunge), Sine loc.: Ovczinnikov 16104 (TAD), EU923786; EU923900; EU661130. Cousinia polytimetica Tschern. (subg. Cousinia
sect. Dichotomae Bunge), Uzbekistan: Bukharskaya reg., deserted left bank of Zeravshan river, to SE from Uzlishkent vil., Kryakin
s.n. (LE), EU923785; EU923899; EU661244. Cousinia pusilla C. Winkl. (subg. Cousinia sect. Dichotomae Bunge), Tajikistan: S
Tajikistan, spackled rock mountains to the S from Besharcha mts. to Babatag range, Botschantzev 117 (LE), EU923787; EU923901;
EU661189. Cousinia auriculata Boiss. (subg. Cousinia sect. Eriocousinia Tschern.), Afghanistan: E, Parvan, Ghowrband, SorkhParsa, 2,050 m, Freitag 1128 (KAS), EU923830; EU923944; EU661191. Cousinia chionophila Rech. & Koie (subg. Cousinia sect.
Eriocousinia Tschern.), Afghanistan: SE, Ghazni, mts. SE of Dashti Nawor (Sperlu Buli), 3,850 m, Freitag 1478 (KAS), EU923829;
EU923943; EU661247. Cousinia gmelini C. Winkl. (subg. Cousinia sect. Eriocousinia Tschern.), Iran: Kandevan to Yoush, Djavadi
s.n. (IRAN), EU923842; EU923956; EU661207. Cousinia polyneura Rech. f. (subg. Cousinia sect. Eriocousinia Tschern.), Afghanistan: E, Kabul, mts. above Paghman, 4,000 m, Freitag 3555 (KAS), EU923781; EU923895; EU661231. Cousinia xanthacantha Regel (subg. Cousinia sect. Eriocousinia Tschern.), Afghanistan: Kabul, in dec. orient. jugi Unai, Breckle 2286 (M), EU923778;
EU923892; EU661230. Cousinia harazensis Rech. f. (subg. Cousinia sect. Harazensis Rech. f.), Iran: Mazandaran, Haraz road,
Rineh, 1,900 m, Mehregan 175 (MJG), EU923825; EU923939; EU661146. Cousinia gatchsaranica I. Mehregan, Assadi & Attar
(subg. Cousinia sect. Haussknechtii Rech. F.), Iran: Gachsaran, N slopes of Khami mt., above Tol Chegah, 2,100 m, Mehregan 83246
(MJG), EU923816; EU923930; EU661159. Cousinia heliantha Bunge (subg. Cousinia sect. Helianthae Bunge), Iran: Khorassan,
30 km from Sabzevar to Esferaien, 1,550 m, Mehregan 170 (MJG), EU923810; EU923924; EU661172. Cousinia coerulea Kult.
ex Tschern. (subg. Cousinia sect. Homalochaete C. Winkl.), Tajikistan: Vorzov canyon, Kudratov, Susanna 2459 & al. (BC),
EU923873; EU923987; EU661144. Cousinia crassipes Kult. (subg. Cousinia sect. Homalochaete C. Winkl.), Iran: Khorassan,
Kalat-e Naderi, Ghareh-Sou, 1,400 m, Mehregan 144 (MJG), EU923811; EU923925; EU661143. Cousinia podophylla Tschern.
(subg. Cousinia sect. Homalochaete C. Winkl.), Sine loc., Ouchinniku 258 (TAD), EU923847; EU923961; EU661233. Cousinia
raddeana C. Winkl. (subg. Cousinia sect. Hoplophylla Tschern.), Iran: Khorassan, Mashhad, Kalat-e Naderi road, Sanganeh, Chahchaheh, 1,600 m, Ghahreman & al. 27315 (TUH), EU923853; EU923967; EU661194. Cousinia immitantiformis Rech.f (subg.
Cousinia sect. Immitantes Rech. f.), Afghanistan: NW, Sabzak pass, NE Herat, 2,400 m, Freitag 6784 (KAS), EU923846; EU923960;
EU661149. Cousinia salangensis Rech. f. (subg. Cousinia sect. Immitantes Rech. f.), Afghanistan: E, Kabul, S Salang valley, marble area, 2,500 m, Freitag 3490 (KAS), EU923845; EU923959; EU661134. Cousinia jassyensis C. Winkl. (subg. Cousinia sect.
Jurineopsis (Juz.) Tschern.), Kyrgyzstan: SW, Jalal Abad Oblast, Toktogul Rayoh, 790 m, Martins 872 (JE), EU923790; EU923904;
EU661240. Cousinia smirnowii Trautv. (subg. Cousinia sect. Kopetdagia Tschern.), Iran: Khorassan, SW Bodjnourd, Salouk mt.,
2,300 m, Mehregan 152 (MJG), EU923802; EU923916; EU661154. Cousinia chrysochlora Rech. f. & Koeie (subg. Cousinia sect.
Lachnosphaerae Rech. f.), Iran: Khorassan, 30 km from Sedeh to Asad Abad, Mahousak, 2,000 m, Mehregan 198 (MJG), EU923856;
EU923970; EU661148. Cousinia lachnosphaera Bunge (subg. Cousinia sect. Lachnosphaerae Rech. f.), Iran: Khorassan, 40 km
from Gonabad to Ferdous, 1,900 m, Mehregan 145 (MJG), EU923857; EU923971; EU661209. Cousinia alepideae Boiss. (subg.
Cousinia sect. Lasiandrae Bunge), Afghanistan: SW, Ghazni, Jamroud, 45 km SW of Ghazni, 2,010 m, Freitag 2008 (KAS),
EU923794; EU923908; EU661182. Cousinia lasiandra Bunge (subg. Cousinia sect. Lasiandrae Bunge), Iran: Khorassan, 3 km
from Doust Abad to Birjand, 1,500 m, Mehregan 127 (MJG), EU923822; EU923936; EU661185. Cousinia triceps Kult. (subg.
Cousinia sect. Leiacanthos Tschern.), Uzbekistan: Kughitang range, Tanghydevol canyon, rocky area, 870 m, Kamelin 870 (LE),
EU923784; EU923898; EU661129. Cousinia antonowii C. Winkl. (subg. Cousinia sect. Leiocaules Bunge), Iran: Khorassan, 10
km from Tivan to Bajgiran, 1,590 m, Mehregan 196 (MJG), EU923849; EU923963; EU661200. Cousinia arctotidifolia Bunge
(subg. Cousinia sect. Leiocaules Bunge), Iran: Khorassan, Shirvan, 10 km from Ziarat to Lojelli, 1,215 m, Mehregan 120 (MJG),
EU923848; EU923962; EU661165. Cousinia heterophylla Boiss. (subg. Cousinia sect. Lepidae Bunge), Afghanistan: W, N-Kattawz,
between Kotanni-Kotal and Patanna, 2,150 m, Freitag 3345 (KAS), EU923804; EU923918; EU661184. Cousinia lepida (Bunge
ex) Boiss. (subg. Cousinia sect. Lepidae Bunge), Iran: Khorassan, Esferaien, 5 km Dahaneh Ojagh to Esferaien, 1,420 m, Mehregan
114 (MJG), EU923805; EU923919; EU661183. Cousinia linczewskii Juz. (subg. Cousinia sect. Leucocaulon Tschern.), Iran: Khorassan, Inter Mashhad and Sarakhs, Mazd-Avand pass, 1,000 m, Mehregan 138 (MJG), EU923850; EU923964; EU661153. Cousinia
turcomanica C. Winkl. (subg. Cousinia sect. Leucocaulon Tschern.), Iran: Khorassan, 110 km W of Bojnourd, between Spakhou
and Kastan, 2,000 m, Mehregan 105 (MJG), EU923834; EU923948; EU661174. Cousinia rigida Kult. (subg. Cousinia sect. Lopholepis Tschern.), Kyrgyzstan: Talasskaya reg., north mountainside of Talassky Alatau, Kur-Bokair canyon, south rocky slope,
Popova s.n. (LE), EU923869; EU923983; EU661222. Cousinia arachnoidea Fisch. & C.A. Mey. (subg. Cousinia sect. Microcarpae Bunge), Kazakhstan: Almatinskaya oblast, Malai Sary pass, 700 m, Susanna 2140 & al. (BC), AY373722*; AY373689*;
EU661248. Cousinia ninae Juz. (subg. Cousinia sect. Microcarpae Bunge), Kyrgyzstan: Oshskaya, Torgulsky reg., Oitaya area
north from Shoporovo vil., speckled soils, Sultanova s.n. (LE), EU923868; EU923982; EU661224. Cousinia platylepis Schrenk.
ex Fisch. & C.A. Mey. (subg. Cousinia sect. Microcarpae Bunge), Kazakhstan: Zambylskaya oblast, 30 km from the Kurdai pass,
Susanna 2158 & al. (BC), AY373737*; AY373704*; EU661236. Cousinia sewertzowii Regel (subg. Cousinia sect. Microcarpae
Bunge), Kazakhstan: Aksu-Dzabagly reservation, 1,800 m, Susanna 2178 & al. (BC), AY373740*; AY373707*; EU661175. Cousinia prolifera Jaub. & Spach (subg. Cousinia sect. Microcousinia Tschern.), Iran: Khorassan, 110 km from Birjand to Ferdous,
1,500 m, Mehregan 233 (MJG), EU923779; EU923893; EU661245. Cousinia heteroloba Rech. f. (subg. Cousinia sect. Molles
Schrenk.), Afghanistan: NE, Andarab valley below Khinjan, near Ghazan, 1,000 m, Freitag 3043 (KAS), EU923776; EU923890;
EU661198. Cousinia candolleana Jaub. & Spach. (subg. Cousinia sect. Myriotemae Rech. f.), Iran: Ghazvin, 10 km from Abgarm
to Avadj, 1,950 m, Mehregan 239 (MJG), EU923858; EU923972; EU661195. Cousinia olgae Regel. & Schmalh. (subg. Cousinia
sect. Olgaeanthe Tschern.), Iran: Khorassan, 40 km from Torbat-e Djam to Saleh Abad, 1,550 m, Mehregan 194 (MJG), EU923780;
EU923894; EU661203. Cousinia eriophylla (Kult.) Bornm. (subg. Cousinia sect. Platyacanthae Rech. f.), Iran: Khorassan, Inter
Ghouchan and Bajgiran, 5 km S of Dorbadam, 1,500 m, Mehregan 190 (MJG), EU923840; EU923954; EU661160. Cousinia rechingerorum Bornm. (subg. Cousinia sect. Platyacanthae Rech. f.), Iran: Khorassan: 30 km W of Torbat-e Djam, E slopes of Bezd mt., 2,000 m, Mehregan 163 (MJG), EU923841; EU923955; EU66116. Cousinia trachyphyllaria Bornm. & Rech. f. (subg. Cousinia sect. Platyacanthae Rech. f.), Iran: Khorassan, Neishabour, Khorw, 2,100 m, Mehregan 217 (MJG), EU923828; EU923942;
EU661163. Cousinia apiculata Tschern. (subg. Cousinia sect. Pseudactinia Tschern.), Iran: Khorassan, Shirvan, Sarani, 2,300 m,
Ghahreman & Attar s.n. (MJG), EU923792; EU923906; EU661162. Cousinia oreodoxa Bornm. & Sint. (subg. Cousinia sect.
Pseudactinia Tschern.), Iran: Khorassan, Inter Bodjnourd and Raz, 7 km from Tangeh Torkeman to Ashkhaneh, 1,000 m, Mehregan
150 (MJG), EU923793; EU923907; EU661151. Cousinia belangeri DC. (subg. Cousinia sect. Pugioniferae Bunge), Iran: Tehran,
15 km to Karaj, 1,200 m, Mehregan s.n. (MJG), EU923819; EU923933; EU661210. Cousinia calcitrapa DC. (subg. Cousinia sect.
Pugioniferae Bunge), Iran: Fars, pass inter Sepidan and Yassoudj, 20 km from Sepidan, 2,400 m, Mehregan 203 (MJG), EU923820;
EU923934; EU661220. Cousinia incompta DC. (subg. Cousinia sect. Pugioniferae Bunge), Iran: 20 km Borujen to Isfahan, 2,400
m, Mehregan 128 (MJG), EU923851; EU923965; EU661186. Cousinia pineticola Rech. f. & Gilli (subg. Cousinia sect. Racemosae
Rech. f.), Afghanistan: E, Kabul, Korogh Koh, SW of Kabul, 2,600 m, Freitag 1568 (KAS), EU923832; EU923946; EU661107.
Cousinia botschantzevii Juz. ex Tschern. (subg. Cousinia sect. Regelianae (Juz.) Tschern.), Uzbekistan: Kuramit mt., Koitash range,
S slope above the Koitash village, 1,850 m, Kamelin 29 (LE), EU923782; EU923896; EU661250. Cousinia autranii C. Winkl.
(subg. Cousinia sect. Rigidissimae Rech. f.), Afghanistan: W, Herat, Ghorat, Bande Frasi, 15 km SSE Farsi, 2,400–2,500 m, Freitag
6831 (KAS), EU923860; EU923974; EU661105. Cousinia rigidissima Rech. f. (subg. Cousinia sect. Rigidissimae Rech. f.), Afghanistan: W, Herat, Ghorat, in summo jugo 5 km S Dahan-e Kaftarkhan ad viam inter Tulak et Farsi, 2,350 m, Podlech 19193 (M),
EU923861; EU923975; EU661187. Cousinia scariosa Regel (subg. Cousinia sect. Scariosae Rech. f.), Afghanistan: N, Sar-e Pol,
Sangcharak, 40 km SSW mts. above Damdaran, 1,900–2,500 m, Freitag 6594 (KAS), EU923838; EU923952; EU661190. Cousinia
eryngioides Boiss. (subg. Cousinia sect. Sciadocousinia Tschern.), Iran: Khorassan, 50 km from Mashhad to Neishabour, 1,500 m,
Mehregan 168 (MJG), EU923836; EU923950; EU661164. Cousinia concolor Bunge (subg. Cousinia sect. Serratuloides Bunge),
Iran: Semnan, Shahroud, S slopes of Shahvar mt., above Nekarmann, 2,500 m, Mehregan 158 (MJG), EU923808; EU923922;
EU661150. Cousinia crispa Jaub. & Spach. (subg. Cousinia sect. Serratuloides Bunge), Iran: Chalous road, 10 km from Kandavan
to Chalous, 2,000 m, Mehregan 185 (MJG), EU923843; EU923957; EU661241. Cousinia hypoleuca Boiss. (subg. Cousinia sect.
Serratuloides Bunge), Iran: Tehran, Inter Firouzkouh and Polur, 5 km W of Lasem, 2,500 m, Mehregan 177 (MJG), EU923807;
EU923921; EU661176. Cousinia pinarocephala Boiss. (subg. Cousinia sect. Serratuloides Bunge), Iran: Tehran, Firouzkouh, Gadouk
pass, 2,200 m, Mehregan 192 (MJG), EU923844; EU923958; EU661223. Cousinia adenosticta Bornm. (subg. Cousinia sect.
Sphaerocephalae Bunge), Iran: Tehran, between Shemshak and Dizin, 3,000 m, Mehregan 165 (MJG), EU923827; EU923941;
EU661110. Cousinia leptolepis (Bornm. & Gauba) Rech. f. (subg. Cousinia sect. Sphaerocephalae Bunge), Iran: Tehran, Karadj,
Kuh Dashteh, S slopes, 2,200 m, Mehregan 111 (MJG), EU923826; EU923940; EU661206. Cousinia shahvarica Rech.f. (subg.
Cousinia sect. Sphaerocephalae Bunge), Iran: Semnan, Shahroud, mt. Shahvar, S slopes, 3,000 m, Mehregan 117 (MJG), EU923823;
EU923937; EU661180. Cousinia xiphiolepis Boiss. (subg. Cousinia sect. Sphaerocephalae Bunge), Iran: Tehran, inter Firouzkouh
and Polur, 5 km W of Lasem, 2,600 m, Mehregan 179 (MJG), EU923824; EU923938; EU661188. Cousinia longifolia C. Winkl.
& Bornm. (subg. Cousinia sect. Spinuliferae Rech. f.), Iran: Kerman, Laleh zar to Rabor, Attar 226 (MJG), EU923815; EU923929;
EU661158. Cousinia qaisarensis Rech. f. (subg. Cousinia sect. Spinuliferae Rech.f.), Afghanistan: NW, Fariab, Maimana, Qaisar,
20 km, mts. above Shakh (Gala Shakh), 2,200 m, Freitag 6745 (KAS), EU923833; EU923947; EU661201. Cousinia alexeenkoana
Bornm. (subg. Cousinia sect. Stenocephalae Bunge), Iran: Isfahan, 10 km from Khaansaar to Boein, 2,500 m, Mehregan 132 (MJG),
EU923799; EU923913; EU661181. Cousinia cylinracea Boiss. (subg. Cousinia sect. Stenocephalae Bunge), Iran: Fars, inter Shiraz
and Sepidan, Shoul pass, 2,000 m, Mehregan 213 (MJG), EU923797; EU923911; EU661108. Cousinia gaubae Bornm. (subg.
Cousinia sect. Stenocephalae Bunge), Iran: Ghazvin, Karadj, in declivibus lapidosis, 1,400 m, Rechinger 5504 (M), EU923818;
EU923932; EU661109. Cousinia tenuiramula Rech. f. (subg. Cousinia sect. Stenocephalae Bunge), Iran: Bakhtiari, Rokh pass,
2,200 m, Mehregan 206 (MJG), EU923798; EU923912; EU661214. Cousinia thamnodes Boiss. & Hausskn. (subg. Cousinia sect.
Stenocephalae Bunge), Iran: Gachsaran, Khami mt. above Tol Chegah, 3,170 m, Mehregan 237 (MJG), EU923800; EU923914;
EU661212. Cousinia komarovii (O. Kuntze) C. Winkl. (subg. Cousinia sect. Stenoloma Juz.), Iran: Khorassan, 15 km Bajgiran to
Ghouchan, 1,700 m, Mehregan 156 (MJG), EU923803; EU923917; EU661152. Cousinia leucantha Bornm. & Sint. (subg. Cousinia sect. Stenoloma Juz.), Iran: Golestan Nat. Park, 12 km ENE of Tange Gol, 1,900–2,100 m, Akhani 11357 (M), EU923837;
EU923951; EU661170. Cousinia stocksii C.Winkl. (subg. Cousinia sect. Stocksianae Rech. f.), Afghanista: W, Farah, Schucht, 1
km nördlich Hakumate Purchaman, 1,880 m, Podlech 21787 (M), EU923795; EU923909; EU661116. Cousinia tenella Fisch. &
C.A. Mey. (subg. Cousinia sect. Tenellae Bunge), Iran: Golestan Nat. Park, between Sharlegh and Cheshmeh Khan, Akhani 243
(MJG), EU923774; EU923888; EU661104. Cousinia chaetocephala Kult. (subg. Cousinia sect. unknown), Iran: Khorassan, Tchenaran, 3km from Meritchegan to Radekan, 1,500 m, Mehregan 115 (MJG), EU923806; EU923920; EU661145. Cousinia pallidivirens Kult. (subg. Cynaroides Tschern. unassigned to any section), Uzbekistan: Sine loc., Botschantzev s.n. (LE), EU923768;
EU923882; EU661126. Cousinia chlorantha Kult. (subg. Cynaroides Tschern. sect. Chrysis Juz.), Uzbekistan: Malguzar mt., S
slope, by Tashkesken ¨say¨, Kamelin 199 (LE), EU923864; EU923978; EU661120. Cousinia karatavica Regel et Schmalh. (subg.
Cynaroides Tschern. sect. Chrysis Juz.), Kazakhstan: Zambylskaya oblast, Karatau mt., Kuyuk pass, 1,000 m, Susanna 2162 & al.
(BC), AY373732*; AY373699*; EU661101. Cousinia korolkowii Regel & Schmalh. (subg. Cynaroides Tschern. sect. Chrysis Juz.),
Uzbekistan: N macro-mountainside, Nuratau range, Sintob kishlak surroundings, rocky slopes, Botschantzev 427 (LE), EU923865;
EU923979; EU661103. Cousinia medians Juz. (subg. Cynaroides Tschern. sect. Chrysis Juz.), Uzbekistan: S, Supkhandarbinskaya
reg., between kishlaks Sajrob and Shurob, 121-122 km of the road from Termez, Speckled rock outcrops, Botschantzev 240 (LE),
EU923769; EU923883; EU661100. Cousinia refracta (Bornm.) Juz. (subg. Cynaroides Tschern. sect. Chrysis Juz.), Tajikistan:
Kondara river canyon, Varzowski Rayon reservation, Susanna 2456 & al. (BC), EU923867; EU923981; EU661111. Cousinia
anomala Franch. (subg. Cynaroides Tschern. sect. Ctenarctium Juz.), Tajikistan: Slopes above Voru, about 300 m above Kishlak,
2,200–2,300 m, Susanna 2521 & al. (BC), EU923770; EU923884; EU661115. Cousinia lappacea Schrenk (subg. Cynaroides
Tschern. sect. Lappaceae Bunge), Kazakhstan: Zambylskaya oblast, Kurdai pass, 900 m, Susanna 2150 & al. (BC), AY373733*; AY373700*; EU661112. Cousinia arctioides Schrenk (subg. Cynaroides Tschern. sect. Nanarctium Juz. ex. Tschern.), Kazakhstan:
Dzhezkazganskaya reg., Turgajskaya lowland, 49 km to SW from Dzhezkazgana, right bank of Kumula river, Tamarix bushland,
Kamelin 6434 (LE), EU923772; EU923886; EU661118. Cousinia triflora Schrenk (subg. Cynaroides Tschern. sect. Oligantha Juz.),
Iran: Golestan Nat. Park, Yakhbala pass, Akhani 102 (MJG), EU923771; EU923885; EU661094. Cousinia albertii Regel et Schmalh.
(subg. Cynaroides Tschern. sect. Pectinatae C. Winkl.), Kazakhstan: Shimkientskaya oblast, Mashat canyon, 1,840 m, Susanna 2206
& al. (BC), AY373721*; AY373688*; EU661099. Cousinia amplissima (Boiss.) Boiss. (subg. Cynaroides Tschern. sect. Pseudarctium Juz.), Iran: Dena, 15 km from Meimand to Yassoudj, 2,500 m, Mehregan 174 (MJG), EU923766; EU923880; EU661098.
Cousinia pseudarctium Bornm. (subg. Cynaroides Tschern. sect. Pseudarctium Juz.), Tajikistan: Vorzov valley, 2 km N kishlak
Ziddy, Susanna 2477 & al. (BC), EU923876; EU923990; EU661095. Cousinia tomentella C. Winkl. (subg. Cynaroides Tschern.
sect. Pseudarctium Juz.), Tajikistan: S mountainside of Guissar range, left bank of Varzob river, Deamalik kishlak surroundings,
1,800 m, Tschukavina 10512 (LE), EU923767; EU923881; EU661097. Cousinia umbrosa Bunge (subg. Cynaroides Tschern. sect.
Pseudarctium Juz.), Kazakhstan: Almatinskaya oblast, Alatau mt. above Almaty, 1,200 m, Susanna 2100 & al. (BC), AY373745*;
AY373712*; EU661096. Cousinia abolinii Kult. ex Tscherneva (subg. Hypacanthodes Tschern. sect. Abolinia Tschern.), Kyrgyzstan:
SW, Jalal Abad Oblast, Kara Saj Tal, Aksy Rayan, 1,030 m, Lazkov s.n. (JE), EU923763; EU923877; EU661113. Cousinia dolichophylla Kult. (subg. Hypacanthodes Tschern. sect. Abolinia Tschern.), Uzbekistan: Ugamsky range above Nanaj, right edge of Pskem
valley, “shiblyak”, Kamelin 265 (LE), EU923875; EU923989; EU661117. Cousinia egregia Juz. (subg. Hypacanthodes Tschern.
sect. Abolinia Tschern.), Uzbekistan: Angren valley, rise to Kamchik pass, rubby slope, Kamelin 420 (LE), EU923866; EU923980;
EU661196. Cousinia grandifolia Kult. (subg. Hypacanthodes Tschern. sect. Amberbopsis Tschern.), Kazakhstan: Zambylskaya
oblast, Talaski Alatau, 6 km W from Il Tai, 1,000 m, Susanna 2181 & al. (BC), AY373730*; AY373697*; EU661114. Cousinia
korshinskyi C. Winkl. (subg. Hypacanthodes Tschern. sect. Lacerae C. Winkl.), Kyrgyzstan: isolated terrain feature Kanka, upper
waters of river Kanka, near snow pot, h. 2,300 m, E.M. Il’ina s.n. (LE), EU923765; EU923879; EU661102. Cousinia macilenta C.
Winkl (subg. Hypacanthodes Tschern. sect. Lacerae C. Winkl.), Tajikistan: SW, Jugum Hissaricum (Gissar), divorticum aquarum
inter flumina Ljuczob et Unou, 3,000 m, Zaprojagaev s.n. (M), EU923764; EU923878; EU661119. |