This draft has not been discussed

S Sambucaceae (incl.)

Comments:

(1) I am in favour of separation of Sambucaceae (Sambucus, Viburnum) as a family, and to place it in sequence closer to Cornaceae and some other families (Diapensiaceae, Hydrangeaceae). (Elven)

WARNING! The family might be split into two.

93.1  Lonicera L. (1753), Sp. Pl. 173.
93.1.1  Lonicera caerulea L. (1753), Sp. Pl. 174.

S

Comments:

A third race strongly approaches the arctic boundary in S Chukotka: subsp. kamtschatica (Sevast.) Gladkova in Jurtz. (1987), Fl. Arct. URSS 10: 28; basionym: L. caerulea L. var. kamtschatica Sevast. in Kraschennikov (1818), Opis. Zemli Kamch. 1: 311. (Elven)

B L. altaica Pall. (1784), Fl. Ross. 1, 1: t. 37.

S

2n=

G RUS SIB

Comments:

(1) Flora Europaea 4: "L. altaica Pallas ..., recorded from N. & E. Russia, is probably only a variant of subsp. (a) [subsp. caerulea] differing in narrower and longer corolla-tube and ellipsoid fruits." (Elven)

B L. pallasii Ledeb. (1821). Ind. Sem. Hort. Dorpat. App. 20.

S

2n=

G RUS SIB

Comments:
93.1.2  Lonicera villosa (Michx.) Roem. & Schult. ***

B

S

B

S

2nD

Comments:

(1) Given by Scoggan (1979) north to Ungava Bay at 5850'N and should perhaps be included as a border case. (Elven)

WARNING! Species and subspecies might be excluded as non-arctic.

93.2  Linnaea L. (1753), Sp. Pl. 631.
93.2.1  Linnaea borealis L. (1753), Sp. Pl. 631.

S

Comments:

S

2n= 32 (4x).

G NOR RUS SIB RFE ALA

Comments:
93.2.1.2  Linnaea borealis L. subsp. americana (Forbes) Hultén (1937), Fl. Aleut. Isl. 310.

B L. americana J. Forbes (1833), Hort. Woburn. 135.

S L. borealis L. var. americana (J. Forbes) Rehd. ***

2n= 32 (4x).

2nD Löve & Löve (1975) list several non-arctic counts.

G ALA CAN GRL

Comments:

(1) Another publication of the varietal name is: var. americana (J. Forbes) Macoun (1912), Geol. Surv. Canad. Summ. Rep. 1911: 25. (Elven)

B Vaccinium album L. (1753), Sp. Pl. 350.

S S. racemosus Michx. (1803), Fl. Bor. Amer. 1: 107; S. rivularis Suksd. (1927), Werdenda 1: 41.

2n=

G CAN?

(1) Subsp. albus may reach or approach the Arctic in the Hudson Bay area acc. to Hultén (1968), but not acc. to Porsild & Cody (1980). Scoggan (1979) does not indicate any clearly arctic site. The western North American subsp. laevigatus (Fern.) Hultén reaches only north to SE Alaska. (Elven)

S

Comments:

B S. pubens Michx. (1803), Fl. Bor. Amer. 1: 181.

S

2n=

G ALA

(1) Hultén (1968) maps 1-2 occurrences just across our arctic boundary in SW Alaska. (Elven)

B V. opulus L. var. (gamma) edule Michx. (1803), Fl. Bor.-Amer. 1: 180.

S

2n=

G RFE ALA CAN

Comments:

94  Adoxaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED

94.1  Adoxa L. (1753), Sp. Pl. 367.
94.1.1  Adoxa moschatellina L. (1753), Sp. Pl. 367.

S

2n= 36 (4x).

G NOR RUS SIB RFE ALA

Comments:

95  Valerianaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED

95.1  Valeriana L. (1753), Sp. Pl. 31.
95.1.1  Valeriana capitata Pall. ex Link (1820), Jahrb. Gewächsk. 1, 3: 66.

S

T [Described from S Siberia: "höchsten Gipfel der Sajanischen Alpen".]

2nD Löve & Löve (1975, 2n=56, nine counts, eight as arctic, 2n=60, four counts, all as arctic); Sokolovskaya & Strelkova (1960, 2n=64); Sokolovskaya (1968, 2n=c.70); Zhukova (1969, 2n=74); Zhukova & Tikhonova (1971, 2n=c.84); Zhukova & Petrovsky (1972, 2n=90, 120).

G RUS SIB RFE ALA CAN

Comments:

S

Comments:

B V. sylvatica Sol. in Banks *** [Frankl. 1st Exp., ed. 2, App. 2]

S V. septentrionalis Rydb. (1900), Mem. New York Bot. Gard. 1: 376.

2n=

G CAN

(1) Valeriana septentrionalis also published by Rydb. in Britt. (1901), Man. Fl. N. U.S. 879. (Elven)

(1) Probably to be treated as an American counterpart (subspecies) of the European   C Asian V. dioica L. s. str. (Elven)
The Valeriana officinalis aggregate (V. officinalis, V. sambucifolia, V. transjenisensis, V. wolgensis)

Comments:

(1) There is probably reasons to keep the diploid(s) apart from the polyploids at species level as alloploidy is most probable. This means that V. officinalis s. str. (2x) should be retained as a species separate from V. sambucifolia (8x). The two Russian entities listed below are much more enigmatic. Valeriana wolgensis is fully included in V. officinalis (subsp. officinalis) by Ockendon (1976, Fl. Eur. 4). I know nothing about V. transjenisensis. Both need some justification if they are to be retained in the checklist. (Elven)
95.1.3  Valeriana officinalis L. (1753), Sp. Pl. 31.

S V. officinalis L. subsp. officinalis.

2n= 14 (2x).

2nD Titz (1964 C Eur); Skalinska (1950 C Eur); Lövkvist & Hultgård (1999 Sweden, 23 counts).

G ICE?

Comments:

S

2n=

G RUS

(1) Probably to be included in V. officinalis. Synonymised with V. officinalis (subsp. officinalis) by Ockendon (1976). (Elven)

S

2n=

G SIB

(1) Reaches, acc. to Fl. Arct. URSS, the Arctic in lower Lena area and probably also along lower Jenisei. Probably to be included in V. officinalis at some level. (Elven)

S V. officinalis L. subsp. sambucifolia (J.C. Mikan ex Pohl) Hayw. (1872), Bot. Pocket Book 70; V. officinalis auct., non L. (1753).

2n= 56 (8x).

2nD Löve & Löve (1975) list numerous counts, one Icelandic; Lövkvist & Hultgård (1999 Sweden, 5 counts).

G NOR RUS

Comments: Subsp. sambucifolia in the Arctic.

(1) The octoploid (2n=56) taxon named as V. salina Pleijel (1925) and often considered a subspecies of V. sambucifolia seems to be mainly Baltic and does not reach the Arctic. Valeriana murmanica Orlova (1966, Fl. Murm. Obl. 5: 426, 175, described from Rybachi Peninsula), which is synonymised with V. salina by Ockendon (1976), is probably only depauperate V. sambucifolia which grows on or near the seashores. The illustration in Fl. Murm. Obl. clearly shows a V. sambucifolia and the description indicates likewise.

There is a recent revision of the Nordic variation (Pedersen unpubl.?) that separates the main variation ('salina' excluded) on two subspecies. I will check what implication this may have on the treatment here. (Elven)

S Lobeliaceae (incl.)

Comments:

(1) Lobeliaceae could well be considered as a separate family. (Elven)

S Syncodon Fourr. (1869), Ann. Soc. Linn. Lyon, n. s., 17: 111 (incl.); Astrocodon Fed. in Kom. (1957), Fl. SSSR 24: 471 (incl.).

Comments:

(1) Syncodon and Astrocodon could perhaps be treated as separate genera as they seem to represent distinct segregates. (Yurtsev & Elven)

S

T [Described from Alaska: Unalaska.]

2nD Löve & Löve (1975) list four counts, one American as arctic.

G RFE ALA

Comments: Subsp. lasiocarpa in the Arctic.

S C. giesekiana auct., non Vest in Roem. & Schult. (1819), Syst. Veg. 5: 89; C. groenlandica Berlin (1884), Öfvers. Kongl. Vet.-Akad. Förh. 41, 7: 50.

2n= (1) 34 (2x). (2) 68 (4x). (3) 102 (6x).

2nD (1-2) Löve & Löve (1975) list numerous counts, many as arctic, for both levels. (1) Croff (1978 N Norw, 11 counts); Lövkvist & Hultgård (1999 S Sweden, 3 counts). (2) Croff (1978 Norw, 126 counts); Lövkvist & Hultgård (1999 S & N Sweden, 29 counts). (3) Kovanda (1970 W Eur).

G ICE NOR RUS SIB RFE ALA* CAN GRL

Comments:

(1) The infraspecific variation is appreciable and treated in different ways in different traditions. Böcher et al. (1978) recognise the Greenland plants as C. gieseckiana with two races, the widely distributed diploid subsp. gieseckiana and the more restricted southern and eastern tetraploid subsp. groenlandica. In Fennoscandia tetraploids occur everywhere except for the northeastern corner. Both lowland plants with many and comparatively small flowers and alpine plants with often large, single flowers are tetraploid. In the northeastern (arctic) corner diploids are dominant or perhaps exclusive (see Croff 1978), with much of the same pattern. Diploids also occur in the southeastern (Swedish and Finnish) lowlands. The few Svalbard plants are diploid and correspond with the Greenland diploid. It seems impossible, at least in the Nordic area, to separate diploids and tetraploids as taxa. (Elven)

(2) A comment has been promised by Yurtsev. (Elven)

S

2n= 34 (2x).

G ICE NOR RUS RFE ALA CAN GRL

Comments:
96.1.4  Campanula tschuktschorum Jurtz. & Fedorov (1974), Bot. Zhurn. 59: 1452.

S

2n=

G RFE

(1) Close to C. uniflora. (Yurtsev & Elven)

S Syncodon glomeratum (L.) Fourr. (1869), Ann. Soc. Linn. Lyon, n. s., 17: 111.

2n= 30 (2x).

2nD Löve & Löve (1975) list eight counts, one Russian as arctic; Lövkvist & Hultgård (1999 Sweden).

G NOR*? RUS* SIB?

Comments:

(1) Established occurrences, probably as garden fugitives, in Rybachi Peninsula and perhaps in arctic N Norway. According to the map of Hultén & Fries (1986) the species reaches the Arctic as native along Jenisei and Lena. (Elven)
96.1.6  Campanula aurita Greene (1888), Pittonia 1: 221.

S Astrocodon auritus (Greene) A.P. Khokhr. (1989), Analiz Fl. Kolymskogo Nagorya 122.

T [Described from Alaska: Yukon R., 63N, leg. Bates.]

2n= 34 (2x).

2nD Knaben (1968 Ala).

G ALA CAN?

Comments:

96.2  Lobelia L. (1753), Sp. Pl. 929.
96.2.1  Lobelia kalmii L. (1753), Sp. Pl. 930.

S

2n=

G CAN