Review of the species of New World Erythroneurini
|
|  Yüklə 1.45 Mb.
|
Type locality: Holotype *, USA, Illinois, Gallatin Co., Gibsonia, on Quercus stellata, 14 VII 1948 (Mills & Ross), (INHS). Distribution: Central and eastern USA. Host plants: Quercus stellata. Notes: E. schista Knull was described based on a specimen with a longer dorsal branch of the pygofer appendage (Fig. 36a2). The relative lengths of the branches vary intraspecifically. 37. Eratoneura longifurca (Hepner, 1966) (Fig. 37, Plate 3p) Erythroneura longifurca Hepner, 1966c:3 Eratoneura longifurca Dietrich & Dmitriev, 2006a:136 Description: Length 2.9–3.1 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes small, narrow, extended dorsomesad. Pygofer lobe rounded; appendage bifurcate far from base, extended to pygofer apex, straight or very slightly curved in dorsal view, curved downward in lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium shorter than shaft; shaft straight and slender in lateral view, compressed, denticulate distally, with lateral lobes at base, without processes; apex acuminate in ventral view. Type locality: Holotype *, USA, Mississippi, Oktibbeha Co., State College, 4 III 1962 (Hepner), (INHS). Distribution: The species is known only from the type locality in Mississippi. Host plants: Unknown. 38. Eratoneura manus (Beamer, 1932) (Fig. 38, Plate 3q) Erythroneura manus Beamer, 1932e:83 Erythroneura tenilla Ross & DeLong, 1950a:294, syn.n. Erythroneura (Eratoneura) manus Young, 1952b:87 Erythroneura colmeri Hepner, 1969a:132, syn.n. Eratoneura manus Dietrich & Dmitriev, 2006a:137 Description: Length 2.9–3.1 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage expanded towards apex, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base, denticulate. Second and third points of style apex very short, toothlike. Dorsal apodeme of aedeagus triangular, without sclerotized connection to anal tube or pygofer appendages; with preatrium about as long as shaft; shaft curved dorsad, broad in lateral view, compressed, denticulate distally, with lateral lobes at base, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Kansas, Leavenworth Co., 28 IV 1928 (Beamer), (KSEM). Distribution: Central and northeastern USA. Host plants: Quercus velutina, Q. stellata, Q. rubra var. ambigua, Q. alba, Q. marilandica, and other species of Quercus. 39. Eratoneura staffordi (Hepner, 1966) (Fig. 39, Plate 3r) Erythroneura staffordi Hepner, 1966c:3 Eratoneura staffordi Dietrich & Dmitriev, 2006a:138 Description: Length 3–3.1 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage bifurcate far from base or expanded towards apex, extended beyond pygofer apex, distinctly sinuate in dorsal view, curved downward distally in lateral view, widest at base, denticulate at apex. Second point of style apex well developed; third point not longer than half distance between other two points; angle between basal and third points about 90°. Aedeagus with preatrium about as long as shaft; shaft curved dorsad, slender in lateral view, compressed, denticulate distally, with lateral lobes at base, without processes; apex acuminate in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Mississippi, Oktibbeha Co., State College, on Carpinus caroliniana, 7 IV 1963 (Hepner), (INHS). Distribution: The species is known only from the type locality in Mississippi. Host plants: Carpinus caroliniana. 40. Eratoneura ardens (McAtee, 1920) (Fig. 40, Plates 1d, 3s) Typhlocyba comes var. scutellaris Gillette, 1898a:764 (prim.hom. of Typhlocyba scutellaris Herrich-Schäffer, 1838c), syn.n. Typhlocyba comes var. scutelleris Gillette, 1898a:764, missp. Erythroneura maculata var. ardens McAtee, 1920a:299 Erythroneura kansana Baker, 1925b:537 n.nov., syn.n. Erythroneura scutellaris var. insolita McAtee, 1926c:133, syn.n. (Plate 3s3) Erythroneura kansana var. ardens Beamer, 1932f:138 Erythroneura ardens Oman, 1949a:95 Erythroneura (Eratoneura) ardens Young, 1952b:86 Eratoneura ardens Dietrich & Dmitriev, 2006a:134 Description: Length 2.7–3.1 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage bifurcate far from base, extended to pygofer apex, distinctly sinuate in dorsal view, straight in lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft curved ventrad, slender in lateral view, round in crossection, denticulate distally, with lateral lobes, without processes; apex blunt in ventral view. Coloration usual for genus, but usually with dark brown mesonotum. Type locality: Holotype *, USA, Maryland, Montgomery Co., Plummers Island, 28 III 1915 (McAtee), (USNM). Distribution: Central and eastern USA, southeastern Canada. Host plants: Ulmus americana, U. alata, and U. rubra. 41. Eratoneura uvaldeana (Knull, 1949) (Fig. 41, Plate 3t) Erythroneura uvaldeana Knull, 1949a:125 Erythroneura (Eratoneura) uvaldeana Young, 1952b:88 Eratoneura uvaldeana Dietrich & Dmitriev, 2006a:139 Description: Length 2.6–2.9 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, extended beyond 3S posterior margin. Pygofer lobe rounded; appendage compressed, bifurcate far from base, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base, denticulate. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft curved dorsad, slender in lateral view, round in crossection, denticulate distally, with lateral lobes at base, without processes; apex blunt in ventral view. Forewings with oblique vittae forming continuous zigzag pattern, without crossbands. Type locality: Holotype *, USA, Texas, Uvalde Co., Uvalde, Garner Park, on Quercus sp., 4 VIII 1937 (Knull), (OSU). Distribution: Texas. Host plants: Quercus sp. 42. Eratoneura inepta (Beamer, 1932) (Fig. 42, Plate 4a) Erythroneura inepta Beamer, 1932g:162 Erythroneura (Eratoneura) inepta Young, 1952b:87 Erythroneura hutchinsi Hepner, 1966c:2, syn.n. (Plate 4a2) Eratoneura inepta Dietrich & Dmitriev, 2006a:136 Description: Length 3.1–3.3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage compressed, bifurcate far from base, extended to pygofer apex, compressed, distinctly sinuate in dorsal view, curved downward in lateral view, widest at middle, denticulate dorsally. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, round in crossection, denticulate distally, with lateral lobes at base, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Kansas, Anderson Co., 26 XI 1927 (Beamer), (KSEM). Distribution: Central and eastern USA. Host plants: Quercus michauxii and Q. stellata. 43. Eratoneura firma (Beamer, 1932) (Fig. 43, Plate 4b) Erythroneura firma Beamer, 1932a:12 Erythroneura (Eratoneura) firma Young, 1952b:87 Erythroneura velox Ross, 1953b:190, syn.n. Erythroneura thaxtoni Hepner, 1966c:2, syn.n. Erythroneura cunninghami Hepner, 1966c:3, syn.n. Erythroneura shumiquera Hepner, 1972a:431, syn.n. Erythroneura combesi Hepner, 1972c:267, syn.n. Eratoneura firma Dietrich & Dmitriev, 2006a:135 Description: Length 2.9–3.3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage bifurcate far from base, extended to pygofer apex, thickened along dorsal margin, distinctly sinuate in dorsal view, curved downward in lateral view, widest at middle. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft curved ventrad, slender in lateral view, round in crossection, denticulate distally, with lateral lobes at base, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Kansas, Anderson Co., 9 IX 1927 (Beamer), (KSEM). Distribution: Central and eastern USA. Host plants: Quercus lyrata, Q. michauxii, Q. stellata, Q. prinus, and other species of Quercus. Notes: Taxa here treated as synonyms appear to represent intraspecific variation in the shape and relative length of the branches of the pygofer appendage (Fig. 43a). 44. Eratoneura bispinosa (Beamer, 1931) (Fig. 44, Plate 4c) Erythroneura bispinosa Beamer, 1931b:241 Erythroneura (Eratoneura) bispinosa Young, 1952b:86 Eratoneura bispinosa Dietrich & Dmitriev, 2006a:134 Description: Length 2.9–3.2 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, extended beyond 3S posterior margin. Pygofer lobe rounded; appendage strongly compressed, bifurcate far from base, extended beyond pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at middle. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft straight and broad in lateral view, round in crossection, denticulate distally, with lateral lobes at base, without processes; apex truncate in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Kansas, Anderson Co., 9 IX 1927 (Beamer), (KSEM). Distribution: Central and eastern USA. Host plants: Ulmus americana, U. alata, U. rubra, Ilex decidua. 45. Eratoneura hyalina (Knull & Auten, 1937) (Fig. 45, Plate 4d) Erythroneura hyalina Knull & Auten, 1937a:575 Erythroneura (Eratoneura) hyalina Young, 1952b:87 Eratoneura hyalina Dietrich & Dmitriev, 2006a:136 Description: Length 2.8 mm. Forewing outer apical cell elongate, more than 2X as long as wide. Pygofer lobe rounded; appendage bifurcate far from base, with ventral branch considerably longer than dorsal, extended to pygofer apex, curved downward in lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, denticulate distally, with lateral lobes, without processes; apex truncate in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Georgia, DeKalb Co., Decatur, 17 IV 1934 (Auten), (OSU). Distribution: The species is known only from the type locality in Georgia. Host plants: Unknown. 46. Eratoneura harnedi (Hepner, 1966) (Fig. 46, Plate 4e) Erythroneura harnedi Hepner, 1966c:5 Eratoneura harnedi Dietrich & Dmitriev, 2006a:136 Description: Length 2.7–2.8 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage bifurcate far from base or three pointed, extended to pygofer apex, distinctly sinuate in dorsal and lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points about 90°. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, round in crossection, denticulate distally, with lateral lobes, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Illinois, Gallatin Co., Shawneetown, on Quercus imbricaria, 14 VII 1948 (Mills & Ross), (INHS). Distribution: Illinois. Host plants: Quercus imbricaria. 47. Eratoneura unica (Beamer, 1932) (Fig. 47, Plate 4f) Erythroneura unica Beamer, 1932e:83 Erythroneura quercalbae Ross & DeLong, 1950a:294, syn.n. Erythroneura (Eratoneura) unica Young, 1952b:88 Eratoneura unica Dietrich & Dmitriev, 2006a:139 Description: Length 2.8–3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes small, narrow, extended dorsomesad. Pygofer lobe rounded; appendage bifurcate far from base, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base. Second and third points of style apex very short, toothlike. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, compressed, denticulate distally, with lateral lobes, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Iowa, Clayton Co., 19 IV 1930 (Beamer), (KSEM). Distribution: Central and northeastern USA. Host plants: Quercus alba. 48. Eratoneura mira (Beamer, 1932) (Fig. 48, Plate 4g) Erythroneura mira Beamer, 1932c:45 Erythroneura (Eratoneura) mira Young, 1952b:87 Eratoneura mira Dietrich & Dmitriev, 2006a:137 Description: Length 2.7–3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes small, narrow, extended dorsomesad. Pygofer lobe rounded; appendage bifurcate far from base, not extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, compressed, denticulate distally, with lateral lobes at base, without processes; apex acuminate in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Illinois, Wabash Co., 31 III 1929 (Beamer), (KSEM). Distribution: Central and eastern USA, two specimens are known from Colorado. Host plants: Quercus alba. 49. Eratoneura facota (Beamer, 1932) (Fig. 49, Plate 4h) Erythroneura facota Beamer, 1932d:70 Erythroneura facata DeLong & Caldwell, 1937c:77, missp. Erythroneura (Eratoneura) facota Young, 1952b:87 Eratoneura facota Dietrich & Dmitriev, 2006a:135 Description: Length 2.6–2.8 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage bifurcate far from base, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base. Second point of style apex very short, toothlike; third point not longer than half distance between other two points; angle between basal and third points less than 90°. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, round in crossection, denticulate distally, with lateral lobes, without processes; apex blunt in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Florida, Pasco Co., Lacoochee, on Quercus sp., 18 VIII 1930 (Beamer), (KSEM). Distribution: Florida. Host plants: Quercus sp. 50. Eratoneura cristata (Knull, 1951) (Fig. 50, Plate 4i) Erythroneura cristata Knull, 1951b:174 Erythroneura (Eratoneura) cristata Young, 1952b:120 Erythroneura spiniterma Hepner, 1969a:133, syn.n. Eratoneura cristata Dietrich & Dmitriev, 2006a:135 Description: Length 2.6–2.8 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage expanded towards apex, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at middle, denticulate apically. Second and third points of style apex very short, toothlike. Aedeagus with preatrium about as long as shaft; shaft straight and slender in lateral view, compressed, denticulate distally, without lateral lobes or processes; apex acuminate in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Ohio, Scioto Co., 10 VI 1944 (Knull), (OSU). Distribution: Central USA. Host plants: Quercus alba, Q. stellata. 51. Eratoneura mcateei sp.n. (Fig. 51, Plate 4j) Description: Length 2.8–3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes large, broad, reaching 3S posterior margin. Pygofer lobe rounded; appendage expanded to apex, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base, denticulate apically. Second and third points of style apex very short, toothlike. Dorsal apodeme of aedeagus parallelsided, connection to pygofer membranous; preatrium about as long as shaft; shaft curved dorsad, broad in lateral view, round in crossection, denticulate distally, with lateral lobes at base, without processes; apex blunt in ventral view. Dorsum yellow or white with orange color pattern; vertex with orange parallel submedial lines with lateral branch, midline pale; anteclypeus pale, concolorous with rest of face; pronotum with Y-shaped medial vitta; mesonotum pale, with orange lateral triangles and apex; thoracic venter entirely pale. Forewings with broken oblique vittae, without crossbands; clavus with separate basal and distal vittae; dark spot at costal margin; inner apical cell with brown spot basally. Diagnosis: Similar to E. manus (Beamer), but with the aedeagal shaft less curved in lateral view and with large lateral lobes at the base of the shaft. Type locality: Holotype *, USA, Illinois, Mason Co., Havana, 2 VII 1934 (DeLong & Ross), (INHS). Studied material: Paratypes: USA, 2 *, Illinois, Franklin Co., Christopher, on Quercus marilandica, 5 VIII 1954 (Ross & Moore), (INHS); 2 *, Illinois, Franklin Co., Royalton, on Quercus marilandica, 5 VIII 1954 (Ross & Moore), (INHS); 1 *, Illinois, Henderson Co., Oquawka, sand ridge, on Quercus sp., 3 VII 1934 (DeLong & Ross), (INHS); 75 *, Illinois, Mason Co., Forest City, on Quercus marilandica, 11 IX 1953 (Stannard & Ross), (INHS); 1 *, Illinois, Mason Co., Forest City, on Quercus marilandica, 11 IX 1953 (Sanderson & Ross), (INHS); 23 *, Illinois, Mason Co., Forest City, on Quercus marilandica, 8 IX 1954 (Ross & Stannard), (INHS); 2 *, Illinois, Mason Co., Havana, 2 VII 1934 (DeLong & Ross), (INHS); 1 *, Illinois, Mason Co., Mason County State Wildlife Refuge, on Quercus sp., 21 V 1953 (Moore), (INHS); 1 *, Illinois, Morgan Co., Meredosia, on Quercus marilandica, 2 X 1951 (Sanderson & Stannard), (INHS); 32 *, Illinois, Morgan Co., Meredosia, on Quercus marilandica, 8 IX 1954 (Ross & Stannard), (INHS). Other studied material from Kansas, Illinois, Mississippi, and Maryland excluded from paratypes. Distribution: Central and norteastern USA. Host plants: Quercus marilandica. Notes: The species is named in honor of Dr. W.L. McAtee, a researcher who described many species of Erythroneura at the beginning of the 20th century. 52. Eratoneura ungulata (Beamer, 1932) (Fig. 52, Plate 4k) Erythroneura ungulata Beamer, 1932d:69 Erythroneura (Eratoneura) ungulata Young, 1952b:88 Eratoneura ungulata Dietrich & Dmitriev, 2006a:139 Description: Length 2.8–3 mm. Forewing outer apical cell elongate, more than 2X as long as wide. 2S abdominal apodemes small, narrow, extended dorsomesad. Pygofer lobe rounded; appendage expanded towards apex, extended to pygofer apex, distinctly sinuate in dorsal view, curved downward in lateral view, widest at base, denticulate at apex. Second and third points of style apex very short, toothlike. Aedeagus with preatrium longer than shaft; shaft curved dorsad, slender in lateral view, compressed, denticulate distally, without lateral lobes or processes; apex acute in ventral view. Coloration usual for genus. Type locality: Holotype *, USA, Louisiana, Natchitoches Co., Natchitoche Parish, 16 VIII 1928 (Beamer), (KSEM). Distribution: South of central and southeastern USA. Host plants: Quercus stellata. 53. Eratoneura distincta (Knull & Auten, 1937) (Fig. 53, Plate 4l) Erythroneura distincta Knull & Auten, 1937a:572 Erythroneura (Eratoneura) distincta Young, 1952b:88 Erythroneura priniquera Hepner, 1967b:68, syn.n. Eratoneura distincta Dietrich & Dmitriev, 2006a:135 |