References: Silva-Brum, 1985: 55; Young, 1991: 194.
Geographic distribution: Pacific and Atlantic. Brazil (RN).
Genus Xenobalanus Steenstrup, 1852
Xenobalanus globicipitis Steenstrup, 1852
Synonymy: Cornwall, 1927: 510.
Description and diagnosis: Darwin, 1854: 440; Cornwall, 1927: 510; Stubbings, 1965: 902.
References: Young, 1991: 194, fig. 1d-e.
Geographic distribution: Cosmopolitan. Brazil (SP).
Superfamily Tetraclitoidea Gruvel, 1903
Family Tetraclitidae Gruvel, 1903
Subfamily Tetraclitellinae Newman & Ross, 1976
Genus Tetraclitella Hiro, 1939
Tetraclitella divisa (Nilsson-Cantell, 1921)
Synonymy: Ross, 1968: 13.
Description and diagnosis: Nilsson-Cantell, 1921: 362; Stubbings, 1967: 291; Anderson, 1986: 108.
References: Young, 1991: 205, fig. 5a-f; 1995: 234.
Geographic distribution: Circumtropical. Brazil (PI, CE, FN).
Genus Newmanella Ross, 1969
Newmanella radiata (Bruguière, 1789)
Synonymy: Ross, 1969: 242.
Description and diagnosis: Darwin, 1854: 343; Ross, 1969: 242.
References: Lacombe, 1977: 11; Lacombe & Rangel, 1978: 5; Young, 1991: 205, fig. 5g-k; Farrapeira-Assunção et al., 1992b: 25; Young; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (PE, RJ, SP).
Subfamily Tetraclitinae Gruvel, 1903
Genus Tesseropora Pilsbry, 1916
Tesseropora atlantica Newman & Ross, 1977
Synonymy: Newman & Ross, 1977: 208.
Description and diagnosis: Newman & Ross, 1977: 208.
References: Edwards & Lubbock, 1983a: 60; 1983b: 68.
Geographic distribution: Atlantic oceanic islands. Brazil (PP) .
Genus Tetraclita Schumacher, 1817
Tetraclita stalactifera (Lamarck, 1818)
Synonymy: Ross, 1968: 8; Young, 1991: 207.
Description and diagnosis: Darwin, 1854: 329 (as T. porosa communis); Ross, 1968: 8.
References: Darwin, 1854: 329, pl. 10, fig. 1a (as T. porosa communis); Müller, 1864: 59, figs. 55, 61-63 (as T. porosa); 1867: 332, figs. 52-53 (as T. porosa); 1868: 395 (as T. porosa); 1869: 88, figs. 55, 61-63 (as T. porosa); Hoek, 1883: 31 (as T. porosa); Weltner, 1895: 289 (as T. porosa); 1897: 257 (as T. porosa communis); Pilsbry, 1916: 254, pl. 59, figs. 1-5b; Nilsson-Cantell, 1927: 786; Luederwaldt, 1929: 52 (as T. stalactifera floridana); Oliveira, 1940a: 138; 1941: 7, pl. 1, figs. 1-2, pl. 2, fig. 4, pl. 10, figs. 1, 3, 6; Ross, 1968: 8, fig. 2; Lacombe & Monteiro, 1974: 635, figs. 6-7; Lacombe, 1977: 2; Newman & Ross, 1976: 48; Lacombe & Rangel, 1978; Lacombe, 1983: 75; Silva-Brum & Absalão, 1989: 143; Young, 1991: 207, fig. 6a-e; 1995: 234.
Geographic distribution: Western Atlantic. Brazil (from MA to RS).
Tetraclita stalactifera floridana Pilsbry, 1916
Synonymy: Young, 1991: 209.
Description and diagnosis: Pilsbry, 1916: 255.
References: Silva-Brum, 1986: 35 (as T. squamosa rubescens); Young, 1991: 209, fig. 6f-j; 1995: 234.
Geographic distribution: Western Atlantic. Brazil (RJ).
Superfamily Balanoidea Leach, 1817
Family Archaeobalanidae Newman & Ross, 1976
Subfamily Archaeobalaninae Newman & Ross, 1976
Genus Chirona Gray, 1835
Chirona amaryllis (Darwin, 1854)
Synonymy: Darwin, 1854: 279; Hoek, 1913: 179.
Description and diagnosis: Darwin, 1854: 279.
References: Young, 1989a: 212; 1987: 30; Farrapeira-Assunção, 1990: 7; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 1, fig. 1; 1995: 230.
Geographic distribution: Indo-West Pacific and southwestern Atlantic. Brazil (PI, PE, BA).
Comments: Introduced species.
Genus Membranobalanus Hoek, 1813
Membranobalanus declivis (Darwin, 1854)
Synonymy: Zullo & Standing, 1983: 471.
Description and diagnosis: Pilsbry, 1916: 230.
References: Farrapeira-Assunção, 1991b: 118; Young, 1995: 230.
Geographic distribution: Western Atlantic. Brazil (BA).
Genus Conopea Say, 1822
Conopea galeata (Linnaeus, 1771)
Synonymy: Young, 1988: 354.
Description and diagnosis: Darwin, 1854: 220; Pilsbry, 1916: 236; Young, 1988: 354.
References: Young, 1988: 354, figs. 1, 4a; 1994: 3; 1995: 230; 1995: 234.
Geographic distribution: Northeastern Pacific and Western Atlantic. Brazil (from PB to SC).
Subfamily Acastinae Kolbasov, 1993
Genus Acasta Leach, 1817
Acasta cyathus Darwin, 1854
Synonymy: Zullo & Standing, 1983: 471.
Description and diagnosis: Pilsbry, 1916: 244.
References: Farrapeira-Assunção, 1991b: 118; Farrapeira-Assunção et al., 1992b: 25; Young, 1995: 230.
Geographic distribution: Indo-West Pacific and Western Atlantic. Brazil (PE, BA).
Family Pyrgomatidae Gray, 1825
Tribe Pyrgomatini Ross & Newman, 1995
Subfamily Ceratoconchinae Newman & Ross, 1976
Genus Ceratoconcha Kramberger-Gorjanovic, 1889
Ceratoconcha domingensis (Moullins, 1866)
Synonymy: Young & Christoffersen, 1984: 246.
Description and diagnosis: Young & Christoffersen, 1984: 246.
References: Young & Christoffersen, 1984: 246, figs. 2d-f, 4-6; Young, 1989b: 199, 1994: 3; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (from PB to BA).
Ceratoconcha floridana (Pilsbry, 1931)
Synonymy: Young & Christoffersen, 1984: 240.
Description and diagnosis: Young & Christoffersen, 1984: 240.
References: Young & Christoffersen, 1984: 240, figs. 2a-c, 3, 5-6; Young, 1988: 362; 1989b: 199; 1994: 3; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (from RN to RJ, RA).
Ceratoconcha paucicostata Young, 1989
Description and diagnosis: Young, 1989b: 193.
References: Young, 1989b: 193, figs. 1-2; 1994: 3;1995: 230.
Geographic distribution: Western Atlantic. Brazil (BA).
Subfamily Megatrematinae Holthuis, 1982
Genus Megatrema Sowerby, 1825
Megatrema madreporarum (Bosc, 1801)
Synonymy: Young, 1988: 360 .
Description and diagnosis: Young, 1988: 360.
References: Young, 1988: 360, figs. 3, 4c; 1994: 4; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (from PB to BA).
Family Balanidae Leach, 1817
Subfamily Balaninae Leach, 1817
Genus Balanus Da Costa, 1778
Balanus amphitrite group
Balanus amphitrite Darwin, 1854
Synonymy: Henry & McLaughlin, 1975: 30.
Description and diagnosis: Henry & McLaughlin, 1975: 30.
References: Weltner, 1897: 264; Oliveira, 1941: 17, pl. 1, fig. 3, pl. 2, figs. 5-6, pl. 3, fig. 1, pl. 11, figs. 1-5 (as B. amphitrite communis), 21, pl. 4, fig. 4, pl. 5, figs. 1-2, pl. 8, figs. 1-5 (as B. amphitrite fluminensis), 22, pl. 4, fig. 5, pl. 9, figs. 1-4 (as B. amphitrite aeratus); Lacombe & Monteiro, 1974: 636, figs. 8-9; Henry & McLaughlin, 1975: 30, figs. 10-11, 13, pl. 1, pl. 5, fig. g, pl. 9, figs. b-c; Lacombe, 1983: 75; Silva-Brum & Absalão, 1989: 143; Farrapeira-Assunção & Silva, 1991: 132; Farrapeira-Assunção et al., 1992b: 25; Siva-Brum & Martins, 1992: 25; Silva-Brum et al., 1994: 22; Young, 1994: 4, fig. 2; 1995: 230.
Geographic distribution: Cosmopolitan. Brazil (from AP to RS).
Balanus eburneus Gould, 1841
Synonymy: Henry & McLaughlin, 1975: 60.
Description and diagnosis: Henry & McLaughlin, 1975: 60.
References: Luederwaldt, 1919: 399; Oliveira, 1941: 19, pl. 3, figs. 3-6, pl. 4, figs. 2, 7 (as B. amphitrite niveus); Lacombe & Monteiro, 1974: 638, fig. 10; Henry & McLaughlin, 1975: 60, fig. 15, pl. 4, pl. 5, fig. g; Silva-Brum & Absalão, 1989: 143; Farrapeira-Assunção & Silva, 1991: 132; Silva-Brum et al., 1994: 22; Young, 1994: 6, fig. 3; 1995: 234.
Geographic distribution: Cosmopolitan. Brazil (PE, RJ, SP).
Balanus improvisus Darwin, 1854
Synonymy: Henry & McLaughlin, 1975: 68 .
Description and diagnosis: Henry & McLaughlin, 1975: 68.
References: Darwin, 1854: 250, pl. 6, fig. 1a-c; Müller, 1867: 332, pl. 7, figs. 22, 32, pl. 8, figs. 50-51 (as B. improvisus assimilis); 1868: 395, pl. 20, figs. 5-6, 9, 17-18, 24 (as B. improvisus assimilis); Weltner, 1897: 266 (as B. improvisus assimilis); 1898: 5; Borradaile, 1916: 132; Henry, 1974: 976; Lacombe & Monteiro, 1974: 641, fig. 14; Henry & McLaughlin, 1975: 68, fig. 16, pl. 5, fig. a-j; Lacombe, 1977: 4; Lacombe & Rangel, 1978: 5; Silva-Brum & Absalão, 1989: 143; Farrapeira-Assunção & Silva, 1991: 132; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 8, fig. 4; 1995: 230.
Geographic distribution: Cosmopolitan. Brazil (from CE to RS, TI).
Balanus reticulatus Utinomi, 1967
Synonymy: Henry & McLaughlin, 1975: 88.
Description and diagnosis: Henry & McLaughlin, 1975: 89.
References: Farrapeira-Assunção, 1990: 7; Farrapeira-Assunção et al., 1992b: 25; Young, 1995: 230.
Geographic distribution: Circumtropical. Brazil (PE, BA).
Comments: Introduced species.
Balanus subalbidus Henry, 1974
Synonymy: Henry & McLaughlin, 1975: 53.
Description and diagnosis: Henry, 1974: 976; Henry & McLaughlin, 1975: 53.
References: Young, 1994: 10, fig. 5; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (PB).
Balanus venustus Darwin, 1854
Synonymy: Henry & McLaughlin, 1975: 164.
Description and diagnosis: Henry & McLaughlin, 1975: 164.
References: Müller, 1867: 345, pl. 7, figs. 29-31, 33-43, pl. 8, figs. 45, 49 (as hybrid between B. improvisus assimilis and B. armatus); 1868: 405, pl. 20, figs. 4, 8, 11-13, 20, 23 (as hybrid between B. improvisus assimilis and B. armatus); Weltner, 1897: 265 (as B. amphitrite niveus); Pilsbry, 1916: 92, pl. 19, figs. 1-2e (as B. amphitrite niveus); Henry & McLaughlin, 1975: 164, fig. 30, pl. 21, figs. a-f; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 13, fig. 6; 1995: 230.
Geographic distribution: Cosmopolitan. Brazil (from PE to RS).
Balanus trigonus group
Balanus calidus Pilsbry, 1916
Synonymy: Pilsbry, 1916: 118.
Description and diagnosis: Pilsbry, 1916: 118; Zullo, 1966: 235.
References: Young, 1994: 15, fig. 7; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (from AP to MA).
Balanus spongicola Brown, 1844
Synonymy: Pilsbry, 1916: 115.
Description and diagnosis: Pilsbry, 1916: 115.
References: Pilsbry, 1916: 115, pl. 25, figs. 2-4c; Nilsson-Cantell, 1927: 784; Young, 1994: 15, fig. 8; 1995: 230.
Geographic distribution: Indian and Atlantic. Brazil (from SP to RS, "Patros Isle" (?) (Pilsbry, 1916)).
Balanus trigonus Darwin, 1854
Synonymy: Pilsbry, 1916: 111.
Description and diagnosis: Pilsbry, 1916: 111.
References: Müller, 1867: 330, pl. 7, figs. 1-21, 23-28, pl. 8, figs. 44, 46-48, pl. 9, fig. 56 (as B. armatus); 1868: 393, pl. 20, figs. 1-3, 7, 10, 14-16, 19, 21-22 (as B. armatus); Hoek, 1883: 31 (as B. armatus); Weltner, 1895; 289 (as B. armatus); 1897: 262 (as B. armatus); Pilsbry, 1916: 111, pl. 26, figs. 1-13e; Oliveira, 1940a: 138; 1941: 15, pl. 4, fig. 3, pl. 10, figs. 2, 4-5, 7; Lacombe & Monteiro, 1974: 641, fig. 12; Lacombe, 1977: 4; Lacombe & Rangel, 1978: 3; Young, 1988: 357; Silva-Brum & Absalão, 1989: 143; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 17, fig. 9; 1995: 230.
Geographic distribution: Cosmopolitan. Brazil (from AP to RS).
Genus Fistulobalanus Zullo, 1984
Fistulobalanus citerosum (Henry, 1974)
Synonymy: Young, 1994: 19, fig. 10.
Description and diagnosis: Henry, 1974: 976, figs. 4-5; Young, 1994: 19, fig. 10.
References: Oliveira, 1941: 19 (as Balanus amphitrite niveus); Henry, 1974: 976, figs. 4-5; Lacombe & Monteiro, 1974: 638, fig. 11 (as B. pallidus); Henry & McLaughlin, 1975: 123, figs. 23d, e, g, 24, pl. 12; Southward, 1975: 42; Newman & Ross, 1976: 63; Lacombe, 1977: 4 (as B. pallidus); Zullo, 1984: 1330; Silva-Brum & Absalão, 1989: 143; Silva-Brum et al., 1984: 22; Young, 1994: 19, fig. 10; 1995: 230.
Geographic distribution: Southwestern Atlantic. Brazil (from PB to RS).
Subfamily Megabalaninae Newman, 1979
Genus Megabalanus Hoek, 1913
Megabalanus coccopoma (Darwin, 1854)
Synonymy: Henry & McLaughlin, 1986: 25.
Description and diagnosis: Henry & McLaughlin, 1986: 25.
References: Lacombe & Monteiro, 1974: 635, fig. 3; Lacombe & Rangel, 1978: 3; Henry & McLaughlin, 1986: 25, figs. 3h, 7a-7; Young, 1987: 30; Silva-Brum & Absalão, 1989: 143; Young, 1994: 23, fig. 11; 1995: 230.
Geographic distribution: Eastern Pacific and southwestern Atlantic. Brazil (from ES to RS).
Comments: Introduced species.
Megabalanus stultus (Darwin, 1854)
Synonymy: Henry & McLaughlin, 1986: 52; Young, 1988: 357.
Description and diagnosis: Henry & McLaughlin, 1986: 52; Young, 1988: 357.
References: Lacombe & Rangel, 1978: 4, fig. 12; Henry & McLaughlin, 1986: 52, figs. 4m, 13e-h; Young, 1988: 357, figs. 2, 4b; 1994: 26; 1995: 230.
Geographic distribution: Western Atlantic. Brazil (from PB to RJ).
Megabalanus tintinnabulum (Linnaeus, 1758 )
Synonymy: Henry & McLaughlin, 1986: 17.
Description and diagnosis: Henry & McLaughlin, 1986: 17.
References: Müller, 1867: 332; 1868: 395; Weltner, 1897: 260 (as Balanus tintinnabulum communis); Pilsbry, 1916: 63, pl. 13, figs. 1-2e (as B tintinnabulum antillensis); Luederwaldt, 1929: 52; Oliveira, 1940a: 35; 1941: 11, pl. 2, figs. 1-2, pl. 4, fig. 1, pl. 5, fig. 3, pl. 8, fig. 6, 14, pl. 3, fig. 2 (as B. tintinnabulum antillensis); Lacombe & Monteiro, 1974: 635, fig. 4; 641, fig. 13 (as B. tintinnabulum antillensis); Lacombe, 1977: 2, 23 (as M. antillensis); Lacombe & Rangel, 1978: 3, fig. 4; Lacombe, 1983: 75; Eston et al., 1986: 40 (as M. coccopoma); Henry & McLaughlin, 1986: 17, figs. 1e, 2a, g-h, 3a-c, 5a-l; Young, 1987: 30 (as M. antillensis); Holthuis, 1991: 18, fig. 29; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 26, fig. 12; 1995: 230.
Geographic distribution: Cosmopolitan. Brazil (from PI to RS, FN).
Megabalanus vesiculosus Darwin, 1854
Synonymy: Henry & McLaughlin, 1986: 44.
Description and diagnosis: Henry & McLaughlin, 1986: 44.
References: Lacombe & Monteiro, 1974: 635, fig. 1 (as Balanus tintinnabulum var.3 and B. t. spinosus); McLaughlin & Lacombe, 1979: 38, figs. 1-15; Henry & McLaughlin, 1986: 44, figs. 1c, 4g, 11i-l; Farrapeira-Assunção et al., 1992b: 25; Young, 1994: 28, fig. 13; 1995: 230.
Geographic distribution: Southwestern Atlantic. Brazil (PE, RJ, SP, PR, SC).
General Comments: Species cited from Brazil but probably are misidentifications.
Chamaesipho columna (Spengler, 1790): Weltner, 1897: 273 (RJ) (? = Chthamalus bisinuatus).
Balanus glandula Darwin, 1854: Newman & Ross, 1976: 60 (RJ). Locality error (Newman, in litt.).
Balanus laevis Bruguiére, 1789: Weltner, 1897: 263; 1898: 5 (RS).
Balanus perforatus angustus (Gmelin, 1789): Weltner, 1897: 263 (BR).
Pyrgoma anglicum Sowerby, 1823 (=Megatrema anglicum): Lacombe, 1977: 11 (RJ).
References
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