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Gustavo a. S. Melo


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Geographic distribution: Western Atlantic. Brazil (from AL to SP).

Comments: On sand beaches; shallow waters.

Pinnixa chaetopterana Stimpson, 1860

Synonymy: Rathbun, 1918: 151; Schmitt et al., 1973: 104; Williams, 1984: 451.

Description and diagnosis: Righi, 1967: 100; Williams, 1984: 451; Melo, 1996: 430.

References: Rathbun, 1918: 151, pl. 33, figs. 3-6; Righi, 1967: 100, figs. 1-6; Rodrigues da Costa, 1970: 262; Coelho & Ramos, 1972: 197; Powers, 1977: 125; Williams, 1984: 451, fig. 360; Abele & Kim, 1986: 64, fig. 698c-d; Bordin, 1987: 23; Melo et al., 1989: 20; Melo, 1996: 430, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA and Brazil (from PE to RS).

Comments: On mud, shell, and gravel bottoms; polychaete symbionts, also in Callichirus burrows.

Pinnixa cristata Rathbun, 1930

Synonymy: Schmitt et al., 1973: 106; Manning & Felder, 1989: 11.

Description and diagnosis: Manning & Felder, 1989: 11; Melo, 1996: 431.

References: Coelho & Ramos, 1972: 196; Melo, 1996: 431, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Central America, and Brazil (AP, PE, SP).

Comments: On sand and muddy sand bottoms; symbiont of Callianassa and other endofauna.

Pinnixa patagoniensis Rathbun, 1918

Synonymy: Schmitt et al., 1973: 116; Manning & Felder, 1989: 20.

Description and diagnosis: Boschi, 1964: 55; Fennucci, 1975: 175; Manning & Felder, 1989: 20; Melo, 1996: 432.

References: Coelho & Ramos, 1972: 197; Fennucci, 1975: 175, pl. 2c; Melo et al., 1989: 20; Melo, 1990a: 78; Boschi et al., 1992: 79, fig. 92; Melo, 1996: 432, 1 fig.

Geographic distribution: Western Atlantic. Brazil (from RJ to RS).

Comments: On sand beaches, Callichirus or Arenicola symbiont, or free-living.

Pinnixa rapax Bouvier, 1917

Synonymy: Righi, 1967: 105.

Description and diagnosis: Righi, 1967: 105; Boschi et al., 1992: 81; Melo, 1996: 433.

References: Righi, 1967: 105, figs. 15-20; Coelho & Ramos, 1972: 196; Fennucci, 1975: 180; Melo, 1990a: 79; Boschi et al., 1992: 81, fig. 94; Melo, 1996: 433, 1 fig.

Geographic distribution: Western Atlantic. Brazil (from RJ to RS), Uruguay, and Argentina.

Comments: On mud bottoms, from 30 to 70m.

Pinnixa sayana Stimpson, 1860

Synonymy: Schmitt et al., 1973: 119; Williams, 1984: 457.

Description and diagnosis: Righi, 1967: 102; Williams, 1984: 457; Melo, 1996: 434.

References: Coelho & Ramos, 1972: 196; Powers, 1977: 128; Williams, 1984: 457, fig. 367; Abele & Kim, 1986: 65, fig. 696g-h; Bordin, 1987: 23; Melo et al., 1989: 21; Coelho & Coelho-Filho, 1993: 45; Melo, 1996: 434, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA and Brazil (from AP to RS).

Comments: On mud bottoms or in burrows of Arenicola, from shallow waters to 75m.

Genus Dissodactylus Smith, 1870



Dissodactylus crinitichelis Moreira, 1901

Synonymy: Schmitt et al., 1973: 17; Williams, 1984: 438.

Description and diagnosis: Williams, 1984: 438; Melo, 1996: 436.

References: Moreira, 1901: 37; Rathbun, 1918: 119, pl. 27, figs. 1-4 (as D. encopei); Rodrigues da Costa, 1970: 260; Coelho & Ramos, 1972: 196; Fennucci, 1975: 172; Williams, 1984: 438, fig. 439; Abele & Kim, 1986: 64, fig. 688b; Melo et al., 1989: 19; Melo, 1996: 436, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, northern South America, Brazil (from PA to RS), and Argentina.

Comments: On fine sand, coral, and shell bottoms, on echinoids Encope and Clypeaster.

Genus Fabia Dana, 1851



Fabia insularis Melo, 1971

Synonymy: Melo, 1971: 198; Fennucci, 1975: 173 (as F. emiliai).

Description and diagnosis: Melo, 1971: 198; Fennucci, 1975: 173 (as F. emiliai); Melo, 1996: 438 (as F. emiliai).

References: Melo, 1971: 198; Fennucci, 1975: 173, pl. 2a, pl. 3f, m, r (as F. emiliai); Melo et al., 1989: 19 (as F. emiliai); Melo, 1990a: 78 (as F. emiliai); 1996: 438, 1 fig. (as F. emiliai).

Geographic distribution: Western Atlantic. Brazil (from RJ to RS) and Argentina.

Comments: Associated with the bivalves Glycymeris and Anadara.

Fabia sebastianensis Rodrigues da Costa, 1970

Synonymy: Rodrigues da Costa, 1970: 257; Schmitt et al., 1973: 24.

Description and diagnosis: Rodrigues da Costa, 1970: 257; Melo, 1996: 439.

References: Rodrigues da Costa, 1970: 257; Coelho & Ramos, 1972: 195; Melo, 1996: 439, 1 fig.

Geographic distribution: Western Atlantic. Brazil (SP).

Comments: Associated with the bivalve Astrina rigida.

Genus Parapinnixa Holmes, 1894



Parapinnixa bouvieri Rathbun, 1918

Synonymy: Schmitt et al., 1973: 31; Williams, 1984: 447.

Description and diagnosis: Williams, 1984: 447; Melo, 1996: 441.

References: Melo, 1996: 441, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA and Brazil (AP).

Comments: On sand and coral bottoms, among the ventral spines of urchins, from 5 to 75m.
Parapinnixa hendersoni Rathbun, 1918

Synonymy: Schmitt et al., 1973: 32; Williams, 1984: 448.

Description and diagnosis: Williams, 1984: 448; Melo, 1996: 442.

References: Righi, 1967: 100; Coelho & Ramos, 1972: 197; Powers, 1977: 122; Williams, 1984: 448, fig. 358; Abele & Kim, 1986: 64, fig. 692b; Melo et al., 1989: 20; Melo, 1996: 442, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, Venezuela, and Brazil (from MA to ES).

Comments: On sand and coral bottoms, sometimes swimming free, from 40 to 60m.

Genus Tumidotheres Campos, 1989



Tumidotheres maculatus (Say, 1818)

Synonymy: Schmitt et al., 1973: 53 (as Pinnotheres maculatus); Williams, 1984: 441 (as P. maculatus).

Description and diagnosis: Fennucci, 1971: 356 (as P. maculatus); Williams, 1984: 441 (as P. maculatus); Melo, 1996: 443.

References: Rodrigues da Costa, 1970: 255 (as P. maculatus); Coelho & Ramos, 1972: 195 (as P. maculatus); Gouvêa, 1974: 175, figs. 1-4 (as P. maculatus); Fennucci, 1975: 167 (as P. maculatus); Williams, 1984: 441, fig. 353 (as P. maculatus); Abele & Kim, 1986: 65, fig. 700a-b (as P. maculatus); Melo, 1996: 443, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from AL to SP).

Comments: On mud, sand, shell, and gravel bottoms, associated with bivalves and in burrows of Arenicola and Chaetopterus; shallow waters to 50m.

Genus Zaops Rathbun, 1900



Zaops ostreum (Say, 1817)

Synonymy: Schmitt et al., 1973: 61 (as Pinnotheres ostreum); Williams, 1984: 444 (as P. ostreum); Manning, 1993: 528.

Description and diagnosis: Williams, 1984: 444 (as P. ostreum); Melo, 1996: 444.

References: Coelho & Ramos, 1972: 195 (as P. ostreum); Powers, 1977: 124 (as P. ostreum); Williams, 1984: 444, figs. 354-355 (as P. ostreum); Abele & Kim, 1986: 65, fig. 700c-d (as P. ostreum); Manning, 1993: 528, fig. 4; Melo, 1996: 444, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from PE to SC).

Comments: Associated with bivalves Cassostrea, Anomia, Mytilus, and Pecten, and in burrows of polychaetes.

Family Grapsidae MacLeay, 1838

Genus Geograpsus Stimpson, 1858

Geograpsus lividus (H. Milne-Edwards, 1837)

Synonymy: Chace & Hobbs, 1969: 157; Manning & Holthuis, 1981: 226.

Description and diagnosis: Chace & Hobbs, 1969: 157; Melo, 1996: 448.

References: Rathbun, 1918: 234, pl. 55; Chace & Hobbs, 1969: 157, figs. 48, 52a-c; Coelho & Ramos, 1972: 200; Fausto-Filho, 1974: 14; 1975: 83; Powers, 1977: 128; Manning & Holthuis, 1981: 226; Abele & Kim, 1986: 62, fig. 674c; Melo et al., 1989: 22; Melo, 1996: 448, 1 fig.

Geographic distribution: Eastern Pacific, Eastern and Western Atlantic. East coast of USA, Antilles, northen South America, and Brazil (from RJ to RS).

Comments: On mud bottoms, from 70 to 150m.

Genus Goniopsis de Haan, 1883



Goniopsis cruentata (Latreille, 1803)

Synonymy: Chace & Hobbs, 1969: 160.

Description and diagnosis: Chace & Hobbs, 1969: 160; Melo, 1996: 449.

References: Moreira, 1901: 39; Rathbun, 1918: 237, pl. 57; Rodrigues da Costa, 1964: 8; Fausto-Filho, 1966: 34; Chace & Hobbs, 1969: 160; Coelho & Ramos, 1972: 200; Gomes-Corrêa, 1972: 200; Powers, 1977: 128; Abele & Kim, 1986: 62, fig. 674d; Melo et al., 1989: 22; Tavares & Albuquerque, 1989: 104; Melo, 1996: 449.

Geographic distribution: Eastern and Western Atlantic. East coast of USA, Antilles, Guianes, and Brazil (from PA to SC, FN).

Comments: In mangroves, among the roots and on muddy beaches, from supralittoral to intertidal zones.

Genus Grapsus Lamarck, 1801



Grapsus grapsus (Linnaeus, 1758)

Synonymy: Chace & Hobbs, 1969: 163; Manning & Holthuis, 1981: 232.

Description and diagnosis: Chace & Hobbs, 1969: 163; Melo, 1996: 450.

References: Moreira, 1901: 39; Rathbun, 1918: 227, pls. 53-54; Fausto-Filho, 1966: 34; Chace & Hobbs, 1969: 163; Coelho & Ramos, 1972: 201; Fausto-Filho, 1974: 14; Powers, 1977: 129; Holthuis et al., 1980: 40; Manning & Holthuis, 1981: 232; Abele & Kim, 1986: 62, fig. 676a; Melo, 1996: 450, 1 fig.

Geographic distribution: Eastern Pacific and Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from CE to ES, FN, RA, TI).

Comments: In rocky intertidal zones, rock bottoms near the surf zone.

Genus Pachygrapsus Randall, 1840



Pachygrapsus corrugatus (von Martens, 1872)

Synonymy: Rathbun, 1918: 252; Manning & Chace, 1990: 66.

Description and diagnosis: Rathbun, 1918: 252; Manning, 1963: 159; Chace & Hobbs, 1969: 167; Melo, 1996: 452.

References: Chace & Hobbs, 1969: 167; Holthuis et al., 1980: 46; Manning & Chace, 1990: 66, fig. 43; Melo, 1996: 452, 1 fig.

Geographic distribution: Western Atlantic. Antilles and Brazil (PP). Central Atlantic. Ascension Is.

Comments: In tidal pools and on encrusting red algae.
Pachygrapsus gracilis (Saussure, 1858)

Synonymy: Chace & Hobbs, 1969: 167; Manning & Holthuis, 1981: 233.

Description and diagnosis: Chace & Hobbs, 1969: 167; Melo, 1996: 453.

References: Rathbun, 1918: 249, pl. 60, fig. 3, pl. 61, fig. 1; Holthuis, 1959: 239; Chace & Hobbs, 1969: 167, fig. 51; Coelho & Ramos, 1972: 201; Powers, 1977: 130; Abele & Kim, 1986: 62, fig. 668f; Melo et al., 1989: 23; Rodrigues & Brossi-Garcia, 1989: 63, figs. 1-5; Melo, 1996: 453, 1 fig.

Geographic distribution: Eastern Pacific, Eastern Atlantic, Mediterranean Sea, and Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from CE to RS, TI).

Comments: In rocks bottoms, under rocks, and on jetties, occasionally among mangrove roots, in estuaries and reefs.
Pachygrapsus transversus (Gibbes, 1850)

Synonymy: Chace & Hobbs, 1969: 169; Williams, 1984: 459.

Description and diagnosis: Chace & Hobbs, 1969: 169; Williams, 1984: 459; Melo, 1996: 454.

References: Rathbun, 1918: 244, pl. 61, figs. 2-3; Rodrigues da Costa, 1964: 7; Fausto-Filho, 1966: 34; Chace & Hobbs, 1969: 169; Coelho & Ramos, 1972: 201; Gomes-Corrêa, 1972: 11; Fausto-Filho, 1974: 13; Powers, 1977: 130; Manning & Holthuis, 1981: 234; Abele & Kim, 1986: 62, fig. 668g; Melo et al., 1989: 24; Tavares & Albuquerque, 1979: 104; Williams, 1984: 459, fig. 368; Melo, 1996: 454, 1 fig.

Geographic distribution: Eastern Pacific, Eastern Atlantic, Mediterranean Sea, and Western Atlantic. East coast of USA, Antilles, northern coast of South America, Brazil (from CE to RS, TI), and Uruguay.

Comments: In rocky bottoms, intertidal.
Genus Planes Bowdich, 1825

Planes cyaneus Dana, 1851

Synonymy: Chace, 1951: 88.

Description and diagnosis: Chace, 1951: 88; Melo, 1996: 455.

References: Chace, 1951: 65, figs. 1b, 3i-n; 1966: 646; Fausto-Filho, 1968: 44 (as P. minutus); Juanicó, 1976: 145; Manning & Holthuis, 1981: 235; Melo, 1996: 455, 1 fig.

Geographic distribution: Indo-Pacific, Eastern Pacific, Eastern and Western Atlantic. Brazil (from RN to RS), Uruguay, and Argentina.

Comments: Pelagic, on floating objects and algae.

Genus Percnon Gistel, 1848



Percnon gibbesi (H. Milne-Edwards, 1853)

Synonymy: Manning & Holthuis, 1981: 238; Williams, 1984: 462.

Description and diagnosis: Williams, 1984: 462; Melo, 1996: 457.

References: Rathbun, 1918: 337, pl. 105; Coelho & Ramos, 1972: 204 (as P. planissineum); Fausto-Filho, 1974: 15; 1975: 82; Powers, 1977: 131; Manning & Holthuis, 1981: 238; Abele & Kim, 1986: 676b; Williams, 1984: 462, fig. 371; Manning & Chace, 1990: 68; Melo, 1996: 457, 1 fig.

Geographic distribution: Eastern Pacific, Eastern and Western Atlantic. East coast of USA, Antilles, and Brazil (FN).

Comments: On rocky and gravel bottoms, intertidal, also associated with echinoid Diadema.

Genus Plagusia Latreille, 1804



Plagusia depressa (Fabricius, 1775)

Synonymy: Chace & Hobbs, 1969: 169; Manning & Holthuis, 1981: 239; Williams, 1984: 463.

Description and diagnosis: Chace & Hobbs, 1969: 169; Williams, 1984: 463; Melo, 1996: 458.

References: Moreira, 1901: 42; Rathbun, 1918: 332, pl. 101; Coelho & Ramos, 1972: 204; Fausto-Filho, 1974: 15; Powers, 1977: 132; Holthuis et al., 1980: 46; Manning & Holthuis, 1981: 239; Abele & Kim, 1986: 63, fig. 676c; Williams, 1984: 463, fig. 372; Manning & Chace, 1990: 69; Melo, 1996: 458, 1 fig.

Geographic distribution: Eastern and Western Atlantic. East coast of USA, Antilles, and Brazil (from CE to BA, FN, RA, TI, PP).

Comments: On rocky bottoms, in crevices and corals, in tidal pools, intertidal.

Genus Aratus H. Milne-Edwards, 1853



Aratus pisonii (H. Milne-Edwards, 1837)

Synonymy: Chace & Hobbs, 1969: 172.

Description and diagnosis: Chace & Hobbs, 1969: 172; Rodriguez, 1980: 396; Melo, 1996: 339.

References: Rathbun, 1918: 323, pl. 96; Holthuis, 1959: 241; Fausto-Filho, 1966: 35; Chace & Hobbs, 1969: 172, figs. 54, 58a; Coelho & Ramos, 1972: 202; Powers, 1977: 152; Rodriguez, 1980: 396, pl. 63; Abele & Kim, 1986: 62, fig. 674a; Melo et al., 1989: 21; Melo, 1996: 460, 1 fig.

Geographic distribution: Eastern Pacific and Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from PI to SP).

Comments: In estuaries and mangroves, on rocks or jetties.

Genus Armases Abele, 1992



Armases angustipes (Dana, 1852)

Synonymy: Abele, 1992: 40.

Description and diagnosis: Abele, 1992: 40; Melo, 1996: 462.

References: Moreira, 1901: 41 (as Sesarma angustipes); Abele, 1972a: 123 (as S. angustipes); 1972b: 167 (as S. angustipes); Coelho & Ramos, 1972: 203 (as S. angustipes); Powers, 1977: 133 (as S. angustipes); Coelho & Ramos-Porto, 1981: 179 (as S. angustipes); Melo et al., 1989: 24 (as S. angustipes); Abele, 1992: 40; Melo, 1996: 462, 1 fig.

Geographic distribution: Western Atlantic. Mexico, Antilles, and Brazil (from CE to SC).

Comments: Around estuaries, in littoral rocks, and in bromeliads.
Armases benedicti (Rathbun, 1897)

Synonymy: Abele, 1992: 53.

Description and diagnosis: Holthuis, 1959: 248 (as Sesarma (Holometopes) benedicti); Abele, 1992: 53; Melo, 1996: 463.

References: Moreira, 1901: 40 (as S. benedicti); Rathbun, 1918: 316, pl. 93 (as S. benedicti); Holthuis, 1959: 248, fig. 62 (as S. benedicti); Coelho & Ramos, 1972: 203 (as S. benedicti); Abele, 1973: 379 (as S. benedicti); Powers, 1977: 133 (as S. benedicti); Coelho & Ramos-Porto, 1981: 203 (as S. benedicti); Abele & Kim, 1986: 63, fig. 670a (as S. benedicti); Abele, 1992: 53; Melo, 1996: 463, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Venezuela, Guianes, and Brazil (AP, PA).

Comments: On wood and rocks around estuaries, also found associated with the vegetation.

Genus Chasmagnathus de Haan, 1833



Chasmagnathus granulata Dana, 1851

Synonymy: Boschi, 1964: 62.

Description and diagnosis: Boschi, 1964: 62; Boschi et al., 1992: 77; Melo, 1996: 464.

References: Moreira, 1901: 42; Rathbun, 1918: 329, pls. 100-159, fig. 9; Boschi, 1964: 62, pls. 3, 17, 20, 22; Coelho & Ramos, 1972: 202; Melo et al., 1989: 22; Tavares & Albuquerque, 1989: 104; Melo, 1990a: 79; Boschi et al., 1992: 77, fig. 88; Melo, 1996: 464, 1 fig.

Geographic distribution: Western Atlantic. Brazil (from RJ to SC), Uruguay, and Angentina.

Comments: On mud bottoms of estuaries, intertidal, in burrows.
Genus Cyclograpsus H. Milne-Edwards, 1837

Cyclograpsus integer (H. Milne-Edwards, 1837)

Synonymy: Chace & Hobbs, 1969: 173; Manning & Holthuis, 1981: 239.

Description and diagnosis: Chace & Hobbs, 1969: 173; Melo, 1996: 465.

References : Rathbun, 1918: 326, pl. 97, figs. 1-2; Fausto-Filho, 1967: 13; Chace & Hobbs, 1969: 173; Coelho & Ramos, 1972: 202; Fausto-Filho, 1974: 14; Powers, 1977: 132; Manning & Holthuis, 1981: 239; Abele & Kim, 1986: 62, fig. 674b; Melo, 1996: 465, 1 fig.

Geographic distribution: Eastern and Western Atlantic. East coast of USA, Central America, northern South America, and Brazil (from CE to SC).

Comments: Marine brackish regions, in supralittoral and intertidal areas of rock beaches.

Genus Metasesarma H. Milne-Edwards, 1853



Metasesarma rubripes (Rathbun, 1897)

Synonymy: Boschi, 1964: 64; Chace & Hobbs, 1969: 175.

Description and diagnosis: Boschi, 1964: 64; Chace & Hobbs, 1969: 175; Melo, 1996: 466.

References: Rathbun, 1918: 319, pl. 94; Boschi, 1964: 64, pls. 3, 15; Chace & Hobbs, 1969: 175; Coelho & Ramos, 1972: 202; Rodriguez, 1980: 394, pl. 62; Melo et al., 1989: 23; Tavares & Albuquerque, 1989: 104; Boschi et al., 1992: 77, fig. 89; Melo, 1996: 466, 1 fig.

Geographic distribution: Western Atlantic. Central America, northern South America, Brazil (from CE to RS), Uruguay, and Argentina.

Comments: Marine brackish regions, among roots and rock crevices.

Genus Sesarma Say, 1817



Sesarma crassipes Cano, 1889

Synonymy: Abele, 1992: 10.

Description and diagnosis: Abele, 1992: 10; Melo, 1996: 468.

References: Rathbun, 1918: 294; Coelho & Ramos, 1972: 204; Coelho & Ramos-Porto, 1981: 178, fig. 1; Abele, 1992: 10, figs. 4c, 5b, 6-7; Melo, 1996: 468, 1 fig.

Geographic distribution: Western Atlantic. Costa Rica and Brazil (PE, BA).

Comments: In mangrove areas, marine, polyhaline.
Sesarma rectum Randall, 1840

Synonymy: Abele, 1992: 15.

Description and diagnosis: Abele, 1992: 15; Melo, 1996: 469.

References: Rathbun, 1918: 298, pl. 82; Holthuis, 1959: 243, pl. 61; Fausto-Filho, 1966: 34; Chace & Hobbs, 1969: 182, pl. 62; Coelho & Ramos, 1972: 203; Coelho & Ramos-Porto, 1981: 178; Melo et al., 1989: 24; Tavares & Albuquerque, 1989: 106; Abele, 1992: 15, figs. 3b, 4b, 5d, 9; Melo, 1996: 469, 1 fig.

Geographic distribution: Western Atlantic. Northern South America and Brazil (from AP to SC).

Comments: In burrows in the shadows of mangrove trees, euryhaline.

Genus Cyrtograpsus Dana, 1851



Cyrtograpsus affinis (Dana, 1851)

Synonymy: Boschi, 1964: 57.

Description and diagnosis: Boschi, 1964: 57; BoVschi et al., 1992: 77; Melo, 1996: 472.

References: Rathbun, 1918: 264, pl. 67 (as Hemigrapsus affinis); Boschi, 1964: 57, pls. 3, 16; Boschi et al., 1992: 77, fig. 87; Melo, 1996: 472, 1 fig.

Geographic distribution: Eastern Pacific and Western Atlantic. Brazil (from RJ to RS) and Argentina.

Comments: In intertidal pools and under rocks, also in shallow waters.

Cyrtograpsus altimanus Rathbun, 1914

Synonymy: Rathbun, 1918: 262; Boschi, 1964: 60.

Description and diagnosis: Rathbun, 1918: 262; Boschi, 1964: 60; Melo, 1996: 473.

References: Rathbun, 1918: 262, pl. 66; Boschi, 1964: 60, pls. 3, 19; Boschi et al., 1992: 75, fig. 85; Coelho & Ramos, 1972: 202; Bordin, 1987: 22; Melo, 1990a: 80; 1996: 473, 1 fig.

Geographic distribution: Western Atlantic. Brazil (RS), Uruguay, and Argentina.

Comments: Appears to be coastal, among rocks at low tide.

Cyrtograpsus angulatus Dana, 1851

Synonymy: Rathbun, 1918: 261; Boschi, 1964: 58.

Description and diagnosis: Rathbun, 1918: 261; Boschi, 1964: 58; Melo, 1996: 474.
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