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Gustavo a. S. Melo


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Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from AP to RJ).

Comments: On sand, mud, shell, and rock bottoms.
Portunus gibbesii (Stimpson, 1859)

Synonymy: Williams, 1984: 389.

Description and diagnosis: Rathbun, 1930: 49; Williams, 1984: 389; Melo, 1996: 325.

References: Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 325, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Venezuela, Guianes, and Brazil (BA).

Comments: On sand, mud, and shell bottoms, also found in bays and estuaries, from shallow waters to 90m.
Portunus ordwayi (Stimpson, 1860)

Synonymy: Rathbun, 1930: 71; Williams, 1984: 390.

Description and diagnosis: Rathbun, 1930: 71; Williams, 1984: 390; Melo, 1996: 326.

References: Rathbun, 1930: 71, pl. 33; Coelho & Ramos, 1972: 186; Gomes-Corrêa, 1972: 7; Fausto-Filho, 1974: 11; Powers, 1977: 84; Takeda & Okutani, 1983: 147; Williams, 1984: 390, fig. 306; Abele & Kim, 1986: 53, fig. 586b; Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 326, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, Venezuela, Guianes, and Brazil (from AP to RS, FN).

Comments: On sand, shell, and coral bottoms, from shallow waters to 110m.
Portunus rufiremus Holthuis, 1959

Synonymy: Holthuis, 1959: 195.

Description and diagnosis: Holthuis, 1959: 195; Melo, 1996: 327.

References: Coelho & Ramos, 1972: 186; Gomes-Corrêa & Brum, 1980: 61; Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 327, 1 fig.

Geographic distribution: Western Atlantic. Guianes and Brazil (AP, PA).

Comments: On mud, shell, and coral bottoms, from 20 to 45m.

Portunus spinicarpus (Stimpson, 1871)

Synonymy: Rathbun, 1930: 92; Williams, 1984: 392.

Description and diagnosis: Rathbun, 1930: 92; Williams, 1984: 392; Melo, 1996: 328.

References: Rathbun, 1930: 92, pl. 45; Holthuis, 1959: 199; Coelho & Ramos, 1972: 187; Gomes-Corrêa, 1972: 7; Powers, 1977: 85; Takeda & Okutani, 1983: 148; Williams, 1984: 392, fig. 308; Abele & Kim, 1986: 54, fig. 586d; Bordin, 1987: 15; Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 328, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from AP to SC).

Comments: Estuarine, from low salinity waters, also in polluted waters, intertidal to 20m.

Portunus spinimanus Latreille, 1819

Synonymy: Rathbun, 1930: 62; Williams, 1984; 393.

Description and diagnosis: Rathbun, 1930: 62; Williams, 1984; 393; Melo, 1996: 329.

References: Rathbun, 1930: 62, pls. 26-28; Coelho & Ramos, 1972: 186; Powers, 1977: 85; Gomes-Corrêa & Brum, 1980: 61; Williams, 1984: 393, fig. 309; Abele & Kim, 1986: 54, fig. 586a; Bordin, 1987: 16; Melo et al., 1989: 13; Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 329, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from PE to RS).

Comments: In brackish waters of bays and channels, on sand, shell, and mud bottoms, from shallow waters to 90m .

Portunus ventralis (A. Milne-Edwards, 1879)

Synonymy: Rathbun, 1930: 43.

Description and diagnosis: Rathbun, 1930: 43; Melo, 1996: 330.

References: Rathbun, 1930: 43, pl. 13, figs. 1-2; Coelho & Ramos, 1972: 186; Powers, 1977: 86; Abele & Kim, 1986: 54, fig. 584a; Coelho & Ramos-Porto, 1993: 297; Melo, 1996: 330, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, Venezuela, and Brazil (from RN to RJ).

Comments: In intertidal pools and sand beaches, intertidal to 25m.

Genus Scylla de Haan, 1833



Scylla serrata (Forskal, 1775)

Synonymy: Melo, 1983; 159.

Description and diagnosis: Melo, 1983; 159; 1996: 331.

References: Melo, 1983; 159, figs. 1-3; 1996: 331, 1 fig.

Geographic distribution: Indo-Pacific and Western Atlantic. Brazil (SP).

Comments: The single specimen collected in Brazil was on sand bottom at 15m.

Family Geryonidae Colosi, 1924

Genus Chaceon Manning & Holthuis, 1989

Chaceon ramosae Manning, Tavares & Albuquerque, 1989

Synonymy: Manning et al., 1989: 646.

Description and diagnosis: Manning et al., 1989: 646; Melo, 1996: 333.

References: Manning et al., 1989: 646, figs. 1-3; Melo, 1996: 333, 1 fig.

Geographic distribution: Western Atlantic. Brazil (ES, RJ, SP).

Comments: Deep waters, from 600 to 1200m.

Family Xanthidae MacLeay, 1838

Genus Actaea de Haan, 1833

Actaea acantha (H. Milne-Edwards, 1834)

Synonymy: Rathbun, 1930: 261; Boone, 1930: 117.

Description and diagnosis: Rathbun, 1930: 261; Boone, 1930: 117; Takeda & Okutani, 1983: 155; Melo, 1996: 339.

References: Rathbun, 1930: 261, pls. 105-106; Coelho & Ramos, 1972: 189; Fausto-Filho, 1974: 12; 1975: 82; Powers, 1977: 87; Takeda & Okutani, 1983: 155; Abele & Kim, 1986: 55, fig. 614a-b; Bordin, 1987: 19; Melo, 1996: 339, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from AP to PE, FN, RA).

Comments: On sand, shell, and coral bottoms, from shallow waters to 25m.

Genus Allactaea Williams, 1974



Allactaea lithostrota (Williams, 1974)

Synonymy: Williams, 1974b: 19; 1984: 397.

Description and diagnosis: Williams, 1974b: 19; 1984: 397; Melo, 1996: 340.

References: Melo, 1996: 340, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from RJ to RS).

Comments: On sand and coral bottoms, from 50 to 640m.

Genus Banareia A. Milne-Edwards, 1869



Banareia palmeri (Rathbum, 1894)

Synonymy: Rathbun, 1930: 260 (as Actaea palmeri); Boone, 1930: 124 (as A. palmeri).

Description and diagnosis: Rathbun, 1930: 260 (as A. palmeri); Boone, 1930: 124 (as A. palmeri); Melo, 1996: 341.

References: Melo, 1996: 341, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from PA to ES, RA).

Comments: Between sponges and corals, from shallow waters to 150m.

Genus Carpilius Leach, 1823



Carpilius corallinus (Herbst, 1783)

Synonymy: Rathbun, 1930: 240.

Description and diagnosis: Rathbun, 1930: 40; Melo, 1996: 342.

References: Rathbun, 1930: 240, pls. 97-99; Fausto-Filho, 1966: 33; Coelho & Ramos, 1972: 189; Fausto-Filho, 1974: 11; Powers, 1977: 88; Abele & Kim, 1986: 56, fig. 650a; Melo, 1996: 342, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, Colombia, Venezuela, and Brazil (from CE to PE, FN).

Comments: On sand bottoms, coral reefs, and sandstones reefs, intertidal to 20m.

Genus Cataleptodius Guinot, 1968



Cataleptodius floridanus (Gibbes, 1850)

Synonymy: Rathbun, 1930: 297 (as Leptodius floridanus); Manning & Holthuis, 1981: 120.

Description and diagnosis: Rathbun, 1930: 297 (as L. floridanus); Manning & Holthuis, 1981: 120; Melo, 1996: 343.

References: Rathbun, 1930: 297, pl. 137, figs. 1-2 (as L. floridanus); Fausto-Filho, 1968: 44 (as L. floridanus); Coelho & Ramos, 1972: 190; Gomes-Corrêa, 1972: 8; Fausto-Filho, 1974: 12 (as L. floridanus); Powers, 1977: 89; Manning & Holthuis, 1981: 120; Abele & Kim, 1986: 56, fig. 650d-g; Melo, 1996: 343, 1 fig.

Geographic distribution: Eastern and Western Atlantic. East coast of USA, Antilles, Central America, northern South America, and Brazil (from CE to RS, FN, RA).

Comments: On rock and coral bottoms, and in sponges, intertidal to 35m.

Genus Domecia Eydoux & Souleyet, 1842



Domecia acanthophora acanthophora (Desbonne & Schramm, 1867)

Synonymy: Williams, 1984: 417.

Description and diagnosis: Williams, 1984: 417; Melo, 1996: 344.

References: Coelho & Ramos, 1972: 193; Gomes-Corrêa, 1972: 8; Powers, 1977: 89; Williams, 1984: 417, fig. 330; Abele & Kim, 1986: 56, fig. 652b; Manning & Chace, 1990: 54; Melo, 1996: 344, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (PB, PE, PP).

Comments: On rock and coral bottoms and in sponges, intertidal to 145m.

Genus Edwardsium Guinot, 1967



Edwardsium spinimanus (H. Milne-Edwards, 1834)

Synonymy: Rathbun, 1930: 274 (as Medaeus spinimanus).

Description and diagnosis: Rathbun, 1930: 274 (as M. spinimanus); Takeda & Okutani, 1983: 156; Melo, 1996: 345.

References: Coelho & Ramos, 1972: 189; Melo, 1996: 345, 1 fig.

Geographic distribution: Western Atlantic. Antilles, Guianes, and Brazil (from CE to RS).

Comments: On mud bottoms, from 15 to 55m.

Genus Eriphia Latreille, 1817



Eriphia gonagra (Fabricius, 1781)

Synonymy: Rathbun, 1930: 545; Williams, 1984: 419.

Description and diagnosis: Rathbun, 1930: 545; Williams, 1984: 419; Melo, 1996: 346.

References: Rathbun, 1930: 545, pl. 222; Coelho & Ramos, 1972: 192; Fausto-Filho, 1966: 33; Gomes-Corrêa, 1972: 7; Powers, 1977: 90; Williams, 1984: 419, fig. 332; Abele & Kim, 1986: 56, fig. 652c; Melo et al., 1989: 13; Melo, 1996: 346, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Central America, Antilles, northern South America, and Brazil (from PA to SC).

Comments: On rock, coral, and algae bottoms, in sponges, intertidal to 5m.

Genus Euryozius Miers, 1886



Euryozius sanguineus (Linnaeus, 1771)

Synonymy: Manning & Chace, 1990: 55.

Description and diagnosis: Manning & Chace, 1990: 55; Melo, 1996: 347.

References: Holthuis et al., 1980: 38; Manning & Chace, 1990: 55; fig. 36; Melo, 1996: 347, 1 fig.

Geographic distribution: Eastern, Central, and Western Atlantic. Brazil (from PA to CE, PP).

Comments: On sand and gravel bottoms, from 20 to 120m.

Genus Eurypanopeus A. Milne-Edwards, 1880



Eurypanopeus abbreviatus (Stimpson, 1860)

Synonymy: Rathbun, 1930: 404; Williams, 1984: 407.

Description and diagnosis: Rathbun, 1930: 404; Williams, 1984: 407; Melo, 1996: 349.

References: Rathbun, 1930: 404, pl. 172, figs. 1-2; Rodrigues da Costa, 1964: 6; Coelho & Ramos, 1972: 191; Fausto-Filho, 1967: 13; Gomes-Corrêa, 1972: 10; Powers, 1977: 91; Williams, 1984: 407, fig. 322; Abele & Kim, 1986: 56, fig. 616a-b; Melo et al., 1989: 14; Tavares & Albuquerque, 1989: 104; Melo, 1996: 349, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from CE to RS).

Comments: On rock and coral bottoms, intertidal to 5m.
Eurypanopeus dissimilis (Benedict & Rathbun, 1891)

Synonymy: Rathbun, 1930: 411

Description and diagnosis: Rathbun, 1930: 411; Melo, 1996: 350.

References: Rathbun, 1930: 411, pl. 173, figs. 1-2; Coelho & Ramos, 1972: 191; Powers, 1977: 92; Abele & Kim, 1986: 56, fig. 616j; Melo, 1996: 350, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from PE to RS).

Comments: In saline lagoons and bays, shallow waters.
Genus Eurytium Stimpson, 1859

Eurytium limosum (Say, 1818)

Synonymy: Chace & Hobbs, 1969: 153; Williams, 1984: 416.

Description and diagnosis: Chace & Hobbs, 1969: 153; Williams, 1984: 416; Melo, 1996: 351.

References: Rathbun, 1930: 423, pl. 176, figs. 1-2; Holthuis, 1959: 208; Chace & Hobbs, 1969: 153; Coelho & Ramos, 1972: 193; Fausto-Filho, 1967: 13; Gomes-Corrêa, 1972: 8; Powers, 1977: 92; Williams, 1984: 416, fig. 329; Abele & Kim, 1986: 56, fig. 654a-b; Melo et al., 1989: 14; Melo, 1996: 351, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Central America, Antilles, northern South America, and Brazil (from PA to SC).

Comments: In mangroves, mud beaches, and in burrows, intertidal to shallow waters.

Genus Garthiope Guinot, 1990



Garthiope barbadensis (Rathbun, 1921)

Synonymy: Guinot, 1990: 473.

Description and diagnosis: Rathbun, 1930: 446 (as Micropanope barbadensis); Guinot, 1990: 473; Melo, 1996: 353.

References: Rodrigues da Costa, 1964: 6 (as M. barbadensis); Melo, 1996: 353, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (ES).

Comments: On coral bottoms, from shallow waters to 30m.
Garthiope spinipes (A. Milne-Edwards, 1880)

Synonymy: Guinot, 1990: 472.

Description and diagnosis: Rathbun, 1930: 443 (as Micropanope spinipes); Guinot, 1990: 472; Melo, 1996: 354.

References: Rathbun, 1930: 443 (as Micropanope spinipes); Coelho & Ramos, 1972: 192; Gomes-Corrêa, 1972: 10; Powers, 1977: 99; Abele & Kim, 1986: 58, fig. 624f (as Micropanope spinipes); Guinot, 1990: 472, fig. 3a, b, g; Melo, 1996: 354, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Venezuela, and Brazil (from AP to ES).

Comments: On coral reefs and among sponges, intertidal to 60m.

Genus Glyptoxanthus A. Milne-Edwards, 1879



Glyptoxanthus vermiculatus (Lamarck, 1818)

Synonymy: Rathbun, 1930: 266.

Description and diagnosis: Rathbun, 1930: 266; Tavares & Albuquerque, 1990: 67; Melo, 1996: 355.

References: Tavares & Albuquerque, 1990: 67, fig. 2; Melo, 1996: 355, 1 fig.

Geographic distribution: Western Atlantic. Venezuela, Guianes, and Brazil (ES).

Comments: On coral bottoms, from shallow waters to 65m.

Genus Hexapanopeus Rathbun, 1898



Hexapanopeus angustifrons (Benedict & Rathbun, 1891)

Synonymy: Rathbun, 1930: 384; Williams, 1984: 415.

Description and diagnosis: Rathbun, 1930: 384; Williams, 1984: 415; Melo, 1996: 357.

References: Melo, 1996: 357, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from PE to SC).

Comments: On sand, mud, shell, and gravel bottoms, intertidal to 140m.

Hexapanopeus caribbaeus (Stimpson, 1871)

Synonymy: Rathbun, 1930: 399.

Description and diagnosis: Rathbun, 1930: 399; Melo, 1996: 358.

References: Rathbun, 1930: 399, pl. 171, figs. 3-5; Coelho & Ramos, 1972: 191; Abele & Kim, 1986: 57, fig. 618a-c; Coelho & Coelho-Filho, 1993: 40; Melo, 1996: 358, 1 fig.

Geographic distribution: Western Atlantic. Antilles, northern South America, and Brazil (from PA to SC).

Comments: On sand bottoms, intertidal to 55m.

Hexapanopeus paulensis Rathbun, 1930

Synonymy: Williams, 1984: 416.

Description and diagnosis: Rathbun, 1930: 395; Williams, 1984: 416; Melo, 1996: 359.

References: Rathbun, 1930: 395, pl. 170, figs. 5-6; Rodrigues da Costa, 1964: 6; Coelho & Ramos, 1972: 191; Powers, 1977: 94; Williams, 1984: 416, fig. 328; Abele & Kim, 1986: 57, fig. 618e-f; Melo et al., 1989: 15; Melo, 1996: 359, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA and Brazil (from PA to SC).

Comments: On sand, shell, and rock bottoms, intertidal to 5m.
Hexapanopeus schmitti Rathbun, 1930

Synonymy: Rathbun, 1930: 393.

Description and diagnosis: Rathbun, 1930: 393; Melo, 1996: 360.

References: Rathbun, 1930: 393, pl. 169, figs. 3-5; Coelho & Ramos, 1972: 191; Melo et al., 1989: 15; Melo, 1996: 360, 1 fig.

Geographic distribution: Western Atlantic. Brazil (from CE to SC) and Uruguay.

Comments: On sand, mud, and shell bottoms, intertidal to 25m.

Genus Melybia Stimpson, 1871



Melybia thalamita Stimpson, 1871

Synonymy: Rathbun, 1930: 562; Williams, 1984: 430.

Description and diagnosis: Rathbun, 1930: 562; Williams, 1984: 430; Melo, 1996: 361.

References: Rathbun, 1930: 562, pl. 230; Coelho & Ramos, 1972: 194; Powers, 1977: 96; Williams, 1984: 430, fig. 342; Abele & Kim, 1986: 57, fig. 656c; Melo, 1996: 361, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from AP to SP)

Comments: On rock, shell, and coral bottoms, from shallow waters to 200m.

Genus Menippe de Haan, 1833



Menippe nodifrons Stimpson, 1859

Synonymy: Rathbun, 1930: 479; Manning & Holthuis, 1981: 137.

Description and diagnosis: Rathbun, 1930: 479; Melo, 1996: 362.

References: Rathbun, 1930: 479, pl. 198, fig. 3, pl. 199; Holthuis, 1959: 208; Rodrigues da Costa, 1964: 5; Coelho & Ramos, 1972: 192; Gomes-Corrêa, 1972: 9; Fausto-Filho, 1966: 33; Powers, 1977: 97; Manning & Holthuis, 1981: 137; Abele & Kim, 1986: 58, fig. 622b; Melo et al., 1989: 15; Melo, 1996: 362, 1 fig.

Geographic distribution: Eastern and Western Atlantic. East coast of USA, Central America, Antilles, northern South America, and Brazil (from MA to SC).

Comments: In intertidal pools, under rocks and jetties; shallow waters.

Genus Micropanope Stimpson, 1871



Micropanope nuttingi (Rathbun, 1898)

Synonymy: Rathbun, 1930: 450; Williams, 1984: 404.

Description and diagnosis: Rathbun, 1930: 450; Williams, 1984: 404; Melo, 1996: 364.

References: Rathbun, 1930: 450; Coelho & Ramos, 1972: 192; Powers, 1977: 98; Williams, 1984: 404, fig. 318; Abele & Kim, 1986: 58, fig. 626a; Melo, 1996: 364, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from AP to SP).

Comments: On rock, coral, and shell, Porites and Halimeda bottoms, from shallow waters to 180m.

Micropanope pusilla A. Milne-Edwards, 1880

Synonymy: Rathbun, 1930: 431.

Description and diagnosis: Rathbun, 1930: 431; Melo, 1996: 365.

References: Melo, 1996: 365, 1 fig .

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from PA to PB).

Comments: On sand, gravel, shell, and coral bottoms, from 30 to 310m.

Micropanope sculptipes Stimpson, 1871

Synonymy: Rathbun, 1930: 428; Williams, 1984: 405.

Description and diagnosis: Rathbun, 1930: 428; Williams, 1984: 405; Melo, 1996: 366.

References: Melo, 1996: 366, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (from AP to RJ).

Comments: On sand, gravel, and coral bottoms, from 10 to 310m.
Micropanope urinator A. Milne-Edwards, 1881

Synonymy: Rathbun, 1930: 451; Williams, 1984: 405.

Description and diagnosis: Rathbun, 1930: 451; Williams, 1984: 405; Melo, 1996: 367.

References: Melo, 1996: 367, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, and Brazil (PA, MA).

Comments: On sand and coral bottoms, from 150 to 460m.

Genus Panopeus H. Milne-Edwards, 1834



Panopeus americanus Saussure, 1857

Synonymy: Rathbun, 1930: 357.

Description and diagnosis: Rathbun, 1930: 357; Melo, 1996: 369.

References: Rathbun, 1930: 357, pl. 164, figs. 3-4, 6; Coelho & Ramos, 1972: 190; Gomes-Corrêa, 1972: 9; Powers, 1977: 101; Abele & Kim, 1986: 58, fig. 630a-b; Melo et al., 1989: 16; Melo, 1996: 369, 1 fig.

Geographic distribution: Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from MA to SC)

Comments: Under rocks in mud beaches and mangroves, on sand, shell, and mud bottoms, intertidal to 25m.

Panopeus austrobesus Williams, 1983

Synonymy: Williams, 1983: 865.

Description and diagnosis: Williams, 1983: 865; Melo, 1996: 370.

References: Williams, 1983: 865, fig. 1a-b; Melo, 1996: 369, 1 fig.

Geographic distribution: Western Atlantic. Brazil (from RJ to RS).

Comments: In bays, estuaries, and mangroves, in burrows, under rocks and in reefs, intertidal to 30m.

Panopeus bermudensis Benedict & Rathbun, 1891

Synonymy: Rathbun, 1930: 360.

Description and diagnosis: Rathbun, 1930: 360; Melo, 1996: 371.

References: Rathbun, 1930: 360, pl. 165; Fausto-Filho, 1968: 44; Coelho & Ramos, 1972: 191; Gomes-Corrêa, 1972: 10; Powers, 1977: 101; Abele & Kim, 1986: 59, fig. 630c; Melo et al., 1989: 16; Melo, 1996: 371, 1 fig.

Geographic distribution: Eastern Pacific and Western Atlantic. East coast of USA, Antilles, northern South America, and Brazil (from CE to SC).

Comments: In intertidal pools, under rocks, and among oysters, in estuaries and mangroves, on sand, up to 15m.

Panopeus harttii Smith, 1869

Synonymy: Rathbun, 1930: 355; Manning & Chace, 1990: 57.

Description and diagnosis: Rathbun, 1930: 355; Melo, 1996: 372.

References: Rathbun, 1930: 355, pl. 164, figs. 1-2, 5; Coelho & Ramos, 1972: 191; Gomes-Corrêa, 1972: 10; Fausto-Filho, 1974: 13; Powers, 1977: 102; Abele & Kim, 1986: 59, fig. 634a-b; Melo et al., 1989: 17; Manning & Chace, 1990: 57, fig. 37; Melo, 1996: 372, 1 fig.
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