Understory species canopy cover under a range of overstory canopy covers in Native Wet Forest. Overstory includes upper tree, secondary tree, and tree fern canopies combined.
Common Name
|
Scientific Name
|
Understory Species Canopy Cover as a function of Overstory Canopy Cover
|
Overstory Canopy Cover Percent
|
50
|
70
|
90
|
'ohi'a lehua
|
Metrosideros polymorpha (seedlings and saplings)
|
5
|
1
|
tr
|
koa
|
Acacia koa (seedlings and saplings)
|
5
|
1
|
tr
|
mamaki
|
Pipturus albidus
|
5
|
1
|
tr
|
olapa
|
Cheirodendron trigynum
|
10
|
5
|
5
|
State 1, Plant Community 1, Native Wet Forest.
State 2 – Grassland
This state is comprised of three grassland plant communities. Most of the pastures in this ecological site are on former sugar plantations where guineagrass was an agricultural weed. Guineagrass now has taken over these lands as the dominant pasture grass. Kikuyugrass is the dominant grass, sometimes with pangolagrass in some higher elevation areas in Kahuku and Keauhou where these species had been planted. Most of these areas are now being reforested. Some parts of Kapapala Ranch appear to be in a transitional area between guineagrass and kikuyugrass dominance. Information on these kikuyugrass/pangolagrass pastures can be found in Ecological Site Description F161AY500HI, Transition Zone Ohia-Koa-Mamane Forest.
Plant Community 2 (Good Condition Pasture) consists of guineagrass with an admixture of glycine (perennial soybean). Continuous grazing that does not allow the favored forage species time to recover from defoliation results in Plant Community 3 (Poor Condition Pasture), which is dominated by lower value forage species but contains enough remnant guineagrass to allow for a transition back to Plant Community 1 with prescribed grazing. Longer-term continuous grazing leads to Plant Community 4 (Weedy Pasture), which consists of low value grass species and increasing cover of alien shrubs and tree saplings. Improvement of this Plant Community requires weed control and prescribed grazing.
Pathways from this state
To State 1 – Native Forest, via “C&D&M”:
C = weed control; D = lack of fire; D = native plant restoration; M = ungulate exclusion.
It may be possible to recreate a plant community resembling Native Forest from Pasture. Weed control must be applied to pasture species and the many opportunistic plant species that invade the site. Weed control would be a perpetual process to capture and maintain the site at least until a closed canopy of native trees developed. Animal foraging (domestic or feral) would have to be eliminated by excluding all ungulates from the restoration site, but domestic ungulates would be useful to initially reduce grass cover and to manage vegetation outside the restoration site perimeter. Extensive planting of native species would follow. Increased shade from trees growing on the site causes a shift from C4 (warm-season) grass dominance (typically guineagrass or kikuyugrass) to C3 (cool-season) shade-tolerant grasses (typically meadow ricegrass). This meadow ricegrass layer can be very dense and detrimental to establishment of native plants. Attempts have been made to suppress meadow ricegrass by planting native shrubs and tree ferns that produce dense shade near the ground and litter that covers the grass.
To State 3 – Tree Plantation, via “C&G”:
C = weed control; G = exotic timber planting.
Pasture may be converted to Tree Plantation by site preparation and planting of timber species (usually eucalyptus) and weed control.
To State 5 – Weedy Alien Forest, via “K”:
K = abandonment.
Abandonment of pastures leads to rapid invasion of alien tree species that take over from the initial growth of grasses and weedy shrubs. The most common tree species are silk oak, christmasberry, albizia, and common guava.
Plant Community 2 – Good Condition Pasture
The dominant grass species in this pasture type is guineagrass that has volunteered in old sugarcane plantations. In higher elevation areas in Kahuku and Keauhou, kikuyugrass and sometimes pangolagrass have been planted.
Pathways from this plant community
To Plant Community 3, Poor Condition Pasture, via “E”:
E = continuous grazing.
Good Condition Pasture degrades to Poor Condition Pasture by continuous grazing that weakens preferred guineagrass or kikuyugrass and legumes in relation to poor forage species such as Hilograss, narrowleaf carpetgrass, and sedges.
State 2, Plant Community 2, Good Condition Pasture.
This list of plants and their relative proportions are based on near-normal years. Fluctuations in species composition and relative production may change from year to year depending upon precipitation or other climatic factors.
Common/Group Name
|
Scientific Name
|
Symbol
|
Functional Group
|
lbs./acre
|
% Comp
|
GRASSES
|
Naturalized Warm Season Tallgrasses
|
|
1
|
11,200-14,000
|
75-85
|
guineagrass
|
Urochloa maxima
|
URMA3
|
1
|
11,200-14,000
|
75-85
|
Napier elephantgrass
|
Pennisetum purpureum
|
PEPU2
|
1
|
T-140
|
T-1
|
Naturalized Warm Season Mid-Grasses
|
|
2
|
T-700
|
T-10
|
kikuyugrass
|
Pennisetum clandestinum
|
PECL3
|
2
|
T-500
|
T-10
|
pangolagrass
|
Digitaria eriantha
|
DIER
|
2
|
T-200
|
T-5
|
Hilograss
|
Paspalum conjugatum
|
PACO14
|
2
|
T-140
|
T-1
|
Green kyllinga
|
Cyperus brevifolius
|
CYBR
|
2
|
T-140
|
T-1
|
Vaseygrass
|
Paspalum urvillei
|
PAUR2
|
2
|
T-140
|
T-1
|
smutgrass
|
Sporobolus indicus
|
SPIN4
|
2
|
T-140
|
T-1
|
marsh bristlegrass
|
Setaria parviflora
|
SEPA10
|
2
|
T-140
|
T-1
|
broomsedge
|
Andropogon virginicus
|
ANVI
|
2
|
T-140
|
T-1
|
beardgrass
|
Schizachyrium condensatum
|
SCCO10
|
2
|
T-140
|
T-1
|
FORBS
|
Naturalized Forbs
|
|
3
|
140-700
|
1-5
|
glycine / perennial soybean
|
Neonotonia wightii
|
NEWI2
|
3
|
140-420
|
1-3
|
three-flowered tickletrefoil
|
Desmodium triflorum
|
DETR4
|
3
|
140-420
|
1-3
|
lilac tasselflower
|
Emilia sonchifolia
|
EMSO
|
3
|
T-140
|
T-1
|
common sow thistle
|
Sonchus oleraceus
|
SOOL
|
3
|
T-140
|
T-1
|
spiny amaranth
|
Amaranthus spinosus
|
AMSP
|
3
|
T-140
|
T-1
|
SHRUBS
|
Naturalized Shrubs, Half-Shrubs, and Trees
|
|
4
|
140-700
|
1-5
|
sourbush
|
Pluchea carolinensis
|
PLCA10
|
4
|
T-140
|
T-1
|
guava
|
Psidium guajava
|
PSGU
|
4
|
T-140
|
T-1
|
christmasberry
|
Schinus terebinthifolius
|
SCTE
|
4
|
T-140
|
T-1
|
castor bean
|
Ricinum communis
|
RICO3
|
4
|
T-140
|
T-1
|
|
|
|
|
|
|
Annual Production lbs./acre
|
|
|
|
|
|
Above Normal
|
16,000
|
|
|
|
|
Normal
|
14,000
|
|
|
|
|
Below Normal
|
10,000
|
|
|
|
|
Percent Ground Cover
|
|
|
|
|
|
Plant
|
65
|
|
|
|
|
Litter
|
30
|
|
|
|
|
Cryptogams
|
0
|
|
|
|
|
Bare ground
|
5
|
|
|
|
|
State 2, Plant Community 2, Good Condition Pasture.
Plant Community 3 – Poor Condition Pasture
Poor Condition Pasture is dominated by grasses of low forage value such as Hilograss, narrowleaf carpetgrass, and sedges. Desirable forage legumes have been grazed out.
Pathways from this plant community
To Plant Community 2, Good Condition Pasture, via “F”:
F = prescribed grazing.
Poor Condition Pasture can be reconverted to Good Condition Pasture by prescribed grazing. A prescribed grazing plan provides for intensive but temporary grazing of pastures that ensures that cattle consume some low-value forage species along with preferred forages and allows preferred forages time to recover from defoliation. The grazing plan may require splitting the herd, creating additional water sources, and creating multiple pastures by cross-fencing. Invading broomsedge and beardgrass may be controlled by mowing their seed stalks before seed set and by liming to increase soil pH.
To Plant Community 4, Weedy Pasture, via “E”:
E = continuous grazing.
Poor Condition Pasture degrades to Weedy Pasture by long-term continuous grazing. Guineagrass cover is greatly reduced and largely replaced by low-value forage grasses. Weedy forbs such as spiny amaranth, alien blackberries, and alien shrubs such as sourbush have increased. Broomsedge and beardgrass often are the most abundant grass species.
Composite representation of State 2, Plant Community 3, Poor Condition Pasture.
Scientific name
|
%Canopy cover by height class (ft)
|
Total Cover
|
Local
common name
|
NRCS
common name
|
Origin
|
Type
|
NRCS
Code
|
0.1 -
2
|
2.1 -
4.5
|
4.6 -
13
|
13.1 -
40
|
40.1 -
80
|
80.1 -
120
|
Psidium guajava
|
1
|
1
|
1
|
|
|
|
1
|
common guava
|
guava
|
a
|
t
|
PSGU
|
Schefflera actinophylla
|
tr
|
|
|
|
|
|
tr
|
octopus tree
|
octopus tree
|
a
|
t
|
SCAC2
|
Falcataria moluccana
|
tr
|
tr
|
|
|
|
|
tr
|
albizia
|
peacocksplume
|
a
|
t
|
FAMO
|
Schinus terebinthifolius
|
tr
|
1
|
1
|
|
|
|
1
|
christmasberry
|
Brazilian peppertree
|
a
|
t
|
SCTE
|
Grevillea robusta
|
tr
|
tr
|
|
|
|
|
tr
|
silver oak
|
silkoak
|
a
|
t
|
GRRO
|
Pluchea carolinensis
|
tr
|
1
|
1
|
|
|
|
1
|
sourbush
|
cure for all
|
a
|
s
|
PLCA10
|
Rubus argutus
|
tr
|
1
|
|
|
|
|
1
|
Florida blackberry
|
sawtooth blackberry
|
a
|
v
|
RUAR2
|
Rubus ellipticus
|
tr
|
1
|
|
|
|
|
1
|
yellow Himalayan raspberry
|
yellow Himalayan raspberry
|
a
|
v
|
RUEL3
|
Rubus rosifolius
|
tr
|
1
|
|
|
|
|
1
|
thimbleberry
|
West Indian raspberry
|
a
|
v
|
RURO
|
Nephrolepis multiflora
|
1
|
|
|
|
|
|
1
|
scaly swordfern
|
scaly swordfern
|
a
|
f
|
NEHI
|
Pteridium aquilinum
|
tr
|
|
|
|
|
|
tr
|
brackenfern
|
western brackenfern
|
n
|
f
|
PTAQ
|
Dicranopteris linearis
|
tr
|
|
|
|
|
|
tr
|
uluhe
|
Old World forkedfern
|
n
|
f
|
DILI
|
Ageratina riparia
|
tr
|
|
|
|
|
|
tr
|
Hamakua pamakani
|
spreading snakeroot
|
a
|
h
|
AGRI2
|
Elephantopus mollis
|
tr
|
|
|
|
|
|
tr
|
soft elephant'sfoot
|
soft elephant'sfoot
|
a
|
h
|
ELMO5
|
Commelina diffusa
|
1
|
|
|
|
|
|
1
|
honohono
|
climbing dayflower
|
a
|
h
|
CODI5
|
Desmodium sp.
|
tr
|
|
|
|
|
|
tr
|
desmodium
|
ticktrefoil
|
a
|
h
|
DESMO
|
Cirsium vulgare
|
tr
|
tr
|
|
|
|
|
tr
|
bull thistle
|
bull thistle
|
a
|
h
|
CIVU
|
Stachytarpheta dichotoma
|
tr
|
tr
|
|
|
|
|
tr
|
owi
|
branched porterweed
|
a
|
h
|
STDI8
|
Hypochoeris radicata
|
tr
|
|
|
|
|
|
tr
|
hairy catsear
|
hairy catsear
|
a
|
h
|
HYRA3
|
Emilia sonchifolia
|
1
|
|
|
|
|
|
1
|
Flora's paintbrush
|
lilac tasselflower
|
a
|
h
|
EMSO
|
Sonchus oleraceus
|
1
|
|
|
|
|
|
1
|
pualele
|
common sowthistle
|
a
|
h
|
SOOL
|
Amaranthus spinosus
|
1
|
|
|
|
|
|
1
|
spiny amaranth
|
spiny amaranth
|
a
|
h
|
AMSP
|
Rumex acetosella
|
1
|
|
|
|
|
|
1
|
sheep sorrel
|
common sheep sorrel
|
a
|
h
|
RUAC3
|
Polygonum capitatum
|
1
|
|
|
|
|
|
1
|
knotweed
|
pinkhead smartweed
|
a
|
h
|
POCA21
|
Cyperus brevifolius
|
5
|
|
|
|
|
|
5
|
flatsedge
|
flatsedge
|
n,a
|
g
|
CYPER
|
Cyperus polystachyos
|
1
|
|
|
|
|
|
1
|
|
manyspike flatsedge
|
n
|
g
|
CYPO
|
Kyllinga brevifolia
|
5
|
|
|
|
|
|
5
|
|
shortleaf spikesedge
|
a
|
g
|
KYBR
|
Saccharum spontaneum
|
|
|
1
|
|
|
|
1
|
wild sugarcane
|
wild sugarcane
|
a
|
g
|
SASP
|
Axonopus fissifolius
|
20
|
|
|
|
|
|
20
|
narrowleaf carpetgrass
|
common carpetgrass
|
a
|
g
|
AXFI
|
Sporobolus indicus
|
1
|
|
|
|
|
|
1
|
smut grass
|
smut grass
|
a
|
g
|
SPIN4
|
Urochloa maxima
|
|
20
|
|
|
|
|
20
|
guineagrass
|
guineagrass
|
a
|
g
|
URMA3
|
Pennisetum clandestinum
|
10
|
|
|
|
|
|
10
|
kikuyugrass
|
kikuyugrass
|
a
|
g
|
PECL2
|
Anthoxanthum odoratum
|
1
|
|
|
|
|
|
1
|
sweet vernalgrass
|
sweet vernalgrass
|
a
|
g
|
ANOD
|
Andropogon virginicus
|
5
|
|
|
|
|
|
5
|
broomsedge
|
broomsedge bluestem
|
a
|
g
|
ANVI2
|
Schizachyrium condensatum
|
5
|
|
|
|
|
|
5
|
beardgrass
|
Colombian bluestem
|
a
|
g
|
SCCO10
|
Setaria parviflora
|
5
|
|
|
|
|
|
5
|
yellow foxtail
|
marsh bristlegrass
|
a
|
g
|
SEPA10
|
Paspalum conjugatum
|
20
|
|
|
|
|
|
20
|
hilograss
|
hilograss
|
a
|
g
|
PACO14
|
Grasslike
|
80
|
20
|
|
|
|
|
100
|
|
|
|
|
|
Native Forbs
|
|
|
|
|
|
|
|
|
|
|
|
|
Exotic Forbs
|
5
|
|
|
|
|
|
5
|
|
|
|
|
|
Native Vines/Epiphytes
|
|
|
|
|
|
|
|
|
|
|
|
|
Exotic Vines
|
1
|
1
|
|
|
|
|
1
|
|
|
|
|
|
Small ferns
|
1
|
|
|
|
|
|
1
|
|
|
|
|
|
Native Shrubs
|
|
|
|
|
|
|
|
|
|
|
|
|
Exotic Shrubs
|
tr
|
1
|
1
|
|
|
|
1
|
|
|
|
|
|
Native Trees
|
|
|
|
|
|
|
|
|
|
|
|
|
Tree ferns (native)
|
|
|
|
|
|
|
|
|
|
|
|
|
Exotic Trees & tree ferns
|
1
|
1
|
1
|
|
|
|
1
|
|
|
|
|
|
Lichen
|
|
|
|
|
|
|
|
|
|
|
|
|
|